D. L. Achat, M. Fortin, G. Landmann, B. Ringeval, A. et al., Effects of forest management on soil C and N storage: meta 552 analysis, Forest Ecology and Management, vol.140, pp.227-238, 2001.

C. Kaiser, M. R. Kilburn, P. L. Clode, L. Fuchslueger, M. Koranda et al., Exploring the transfer of recent plant photosynthates to soil microbes: mycorrhizal pathway vs 555 direct root exudation, New Phytologist, vol.554, pp.1537-1551, 2015.

H. Kang, S. Kang, and D. Lee, Variations of soil enzyme activities in a temperate forest soil, 2009.

, Ecol. Res, vol.24, pp.1137-1143

Z. Kotroczó, Z. Veres, I. Fekete, Z. Krakomperger, J. A. Tóth et al., Soil enzyme activity in response to long-term organic matter manipulation, Soil Biol, vol.559, 2014.

, Biochem, vol.70, pp.237-243

T. E. Kraus, R. A. Dahlgren, and R. J. Zasoski, Tannins in nutrient dynamics of forest 562 ecosystems-a review, Plant and Soil, vol.256, pp.41-66, 2003.

Y. Kuzyakov, Priming effects: Interactions between living and dead organic matter, Soil Biol, 2010.

, Biochem, vol.42, pp.1363-1371

J. Kyaschenko, K. E. Clemmensen, E. Karltun, and B. D. Lindahl, Below-ground organic 566 matter accumulation along a boreal forest fertility gradient relates to guild interaction within fungal 567 communities, Ecol. Lett, vol.20, pp.1546-1555, 2017.

K. Lajtha, S. E. Crow, Y. Yano, S. S. Kaushal, E. Sulzman et al., , 2005.

, Detrital controls on soil solution N and dissolved organic matter in soils: A field experiment

, Biogeochemistry, vol.76, pp.261-281

K. Lajtha, R. D. Bowden, and K. Nadelhoffer, Litter and root manipulations provide insights 572 into soil organic matter dynamics and stability, Soil Sci. Soc. Am. J, vol.78, p.261, 2014.

R. Lal, Forest soils and carbon sequestration, For. Ecol. Manag, vol.220, pp.242-258, 2005.

J. W. Leff, W. R. Wieder, P. G. Taylor, A. R. Townsend, D. R. Nemergut et al., Experimental litterfall manipulation drives large and rapid changes in soil 576 carbon cycling in a wet tropical forest, Glob. Change Biol, vol.18, pp.2969-2979, 2012.

H. T. Leung, K. R. Maas, R. C. Wilhelm, and W. W. Mohn, Long-term effects of timber 578 harvesting on hemicellulolytic microbial populations in coniferous forest soils, ISME J, vol.10, pp.363-375, 2016.

C. Leuschner, Water extraction by tree fine roots in the forest floor of a temperate Fagus-580 Quercus forest, Ann. Sci. For, vol.55, pp.141-157, 1998.

Q. Li, H. Lee-allen, and A. G. Wollum, Microbial biomass and bacterial functional diversity 582 in forest soils: effects of organic matter removal, compaction, and vegetation control, Soil Biol, 2004.

, Biochem, vol.36, pp.571-579

Y. Liu, G. Wang, K. Yu, P. Li, L. Xiao et al., A new method to optimize root order 585 classification based on the diameter interval of fine root, Sci. Rep, vol.8, 2018.

S. Lladó and P. Baldrian, Community-level physiological profiling analyses show potential to 587 identify the copiotrophic bacteria present in soil environments, PLOS ONE, vol.12, 2017.

K. Lorenz, C. M. Preston, S. Krumrei, and K. H. Feger, Decomposition of needle/leaf litter 589 from Scots pine, black cherry, common oak and European beech at a conurbation forest site, European 590 Journal of Forest Research, vol.123, pp.177-188, 2004.

S. Luyssaert, E. Schulze, A. Börner, A. Knohl, D. Hessenmöller et al., , p.592

J. Grace, Old-growth forests as global carbon sinks, Nature, vol.455, pp.213-215, 2008.
URL : https://hal.archives-ouvertes.fr/cea-00910763

M. C. Fisk and T. J. Fahey, Microbial biomass and nitrogen cycling responses to 594 fertilization and litter removal in young northern hardwood forests, Biogeochemistry, vol.53, pp.201-223, 2001.

K. J. Nadelhoffer, R. D. Boone, R. D. Bowden, J. D. Canary, J. Kaye et al., , p.596

J. A. Aitkenhead, K. Lajtha, and W. H. Mcdowell, The DIRT experiment: litter and root influences 597 on forest soil organic matter stocks and function, Forests in Time: The 598 Environmental Consequences of 1000 Years of Change in New England, pp.300-315, 2004.

J. F. O'hehir and E. K. Nambiar, Productivity of three successive rotations of P. radiata 601 plantations in South Australia over a century, For. Ecol. Manag, vol.259, pp.1857-1869, 2010.

J. Oksanen, F. G. Blanchet, R. Kindt, P. Legendre, P. R. Minchin et al., 603 Community Ecology Package, 2013.

Y. Pan, R. A. Birdsey, J. Fang, R. Houghton, P. E. Kauppi et al., A large and persistent carbon sink in the world's forests, 608 Science, vol.333, pp.988-993, 2011.

P. Perez-batallon, G. Ouro, F. Macias, and A. Merino, Initial mineralization of organic matter in a 610 forest plantation soil following different logging residue management techniques, Ann. For. Sci, vol.58, pp.807-818, 2001.

F. Ponder and M. Tadros, Phospholipid fatty acids in forest soil four years after organic 613 matter removal and soil compaction, Appl. Soil Ecol, vol.19, pp.173-182, 2002.

R. F. Powers, A. Scott, D. Sanchez, F. G. Voldseth, R. A. Page-dumroese et al., The North American long-term soil productivity experiment: Findings from the 616 first decade of research, For. Ecol. Manag, vol.220, pp.31-50, 2005.

J. Preston-mafham, L. Boddy, and P. F. Randerson, Analysis of microbial community 618 functional diversity using sole-carbon-source utilisation profiles -a critique, FEMS Microbiol. Ecol, vol.619, pp.1-14, 2002.

K. Pritsch, P. E. Courty, J. Churin, B. Cloutier-hurteau, M. A. Ali et al., , p.621

S. Egli, J. Ernst, and L. Fraissinet-tachet, Optimized assay and storage conditions for 622 enzyme activity profiling of ectomycorrhizae, Mycorrhiza, vol.21, pp.589-600, 2011.

. R-core-team, R: A language and environment for statistical computing. R Foundation for 624 Statistical Computing, 2016.

E. J. Sayer, Using experimental manipulation to assess the roles of leaf litter in the functioning 627 of forest ecosystems, Biol. Rev, vol.81, pp.1-31, 2006.

R. A. Slesak, B. J. Palik, A. W. Amato, and V. J. Kurth, Changes in soil physical and 629 chemical properties following organic matter removal and compaction: 20-year response of the aspen 630 Lake-States Long Term Soil Productivity installations, For. Ecol. Manag, vol.392, pp.68-77, 2017.

A. Smolander, V. Kitunen, P. Tamminen, and M. Kukkola, Removal of logging residue in 632 Norway spruce thinning stands: Long-term changes in organic layer properties, Soil Biol. Biochem, vol.633, pp.1222-1228, 2010.

D. M. Stone, Sustaining aspen productivity in the Lake States, Sustaining Aspen in Western 635 Landscapes: Symposium Proceedings, pp.13-15, 2000.

D. Shepperd, D. L. Binkley, T. J. Bartos, L. G. Stohlgren, and . Eskew, USDA For. Serv, Proc. RMRS-P-637 118, pp.47-59

T. Wei and V. Simko, R package "corrplot": Visualization of a Correlation Matrix 639 (Version 0.84), 2017.

X. Tan, S. X. Chang, and R. Kabzems, Soil compaction and forest floor removal reduced 641 microbial biomass and enzyme activities in a boreal aspen forest soil, Biol. Fertil. Soils, vol.44, pp.471-479, 2008.

M. A. Vadeboncoeur, S. P. Hamburg, R. D. Yanai, and J. D. Blum, Rates of sustainable forest 643 harvest depend on rotation length and weathering of soil minerals, For. Ecol. Manag, vol.318, pp.194-205, 2014.

Z. Veres, Z. Kotroczó, I. Fekete, J. A. Tóth, K. Lajtha et al., , 2015.

, Soil extracellular enzyme activities are sensitive indicators of detrital inputs and carbon availability

, Appl. Soil Ecol, vol.92, pp.18-23

M. Wang and Q. Zhang, Issues in using the WinRHIZO system to determine physical 648 characteristics of plant fine roots, Acta Ecol. Sin, vol.29, pp.136-138, 2009.

X. Wang, C. Tang, J. Severi, C. R. Butterly, and J. A. Baldock, Rhizosphere priming effect 650 on soil organic carbon decomposition under plant species differing in soil acidification and root 651 exudation, New Phytol, vol.211, pp.864-873, 2016.

K. Wilhelm, B. Rathsack, and J. Bockheim, Effects of timber harvest intensity on 653 macronutrient cycling in oak-dominated stands on sandy soils of northwest Wisconsin, 2013.

. Manag, , vol.291, pp.1-12

R. C. Wilhelm, E. Cardenas, K. R. Maas, H. Leung, L. Mcneil et al., , p.656

J. M. Kranabetter and S. Dubé, Biogeography and organic matter removal shape long-term 657 effects of timber harvesting on forest soil microbial communities, ISME J, vol.11, pp.2552-2568, 2017.

B. Zhang, H. Wang, S. Yao, and L. Bi, Litter quantity confers soil functional resilience through 659 mediating soil biophysical habitat and microbial community structure on an eroded bare land restored 660 with mono Pinus massoniana, Soil Biology and Biochemistry, vol.57, pp.556-567, 2013.

L. ?if?áková, T. V?trovský, A. Howe, and P. Baldrian, Microbial activity in forest soil 662 reflects the changes in ecosystem properties between summer and winter: Seasonal dynamics of a soil 663 microbial community, Environ. Microbiol, vol.18, pp.288-301, 2016.