M. A. Abro, F. Lecompte, F. Bryone, and P. Nicot, Nitrogen fertilization of the host plant influences production and pathogenicity of Botrytis cinerea secondary inoculum, 2012.

A. Adam, Elicitation de la résistance systémique induite chez la tomate et le concombre et activation de la voie de la lipoxygénase par des rhizobactéries non-pathogènes. Mémoire de thèse, 2008.

S. Adamowicz, L. Bot, and J. , preserves the proportionate relation linking long-term organic-nitrogen accumulation to intercepted radiation, New Phytologist, vol.59, issue.3, pp.663-672, 2008.
DOI : 10.2135/cropsci1994.0011183X003400030022x

S. Adamowicz, L. Bot, J. , H. Magana, R. Fabre et al., Totomatix: a novel automatic set-up to control diurnal, diel and long-term plant nitrate nutrition, Annals of Botany, vol.16, issue.1, pp.309-319, 2012.
DOI : 10.1051/agro:19960201

E. A. Ainsworth, A. Rogers, A. D. Leakey, L. E. Heady, Y. Gibon et al., Does elevated atmospheric [CO2] alter diurnal C uptake and the balance of C and N metabolites in growing and fully expanded soybean leaves?, Journal of Experimental Botany, vol.58, issue.3, pp.579-591, 2007.
DOI : 10.1093/jxb/erl233

S. Ajouz, P. C. Nicot, and M. Bardin, Adaptation to pyrrolnitrin in Botrytis cinerea and cost of resistance, Plant Pathology, vol.113, issue.3, pp.556-566, 2010.
DOI : 10.1111/j.1574-6941.2003.tb01042.x

T. Aniszewski, Alkaloids -Secrets of life. Alkaloid chemistry, biological significance, applications and ecological role, 2007.

Y. P. Bajaj, Medicinal and aromatic plants VIII, Biotechnology in Agriculture and Forestry, 2001.
DOI : 10.1007/978-3-662-08612-4

C. J. Baker, B. D. Whitaker, D. P. Roberts, N. M. Mock, C. P. Rice et al., Induction of redox sensitive extracellular phenolics during plant-bacterial interactions". Physiological and Molecular Plant Pathology, pp.90-98, 2005.

I. T. Baldwin, M. J. Karb, and T. E. Ohnmeiss, Allocation of 15 N from nitrate to nicotine: production and turnover of a damage-induced mobile defense, Ecology, issue.6, pp.75-1703, 1994.

E. K. Barto and D. Cipollini, Testing the optimal defense theory and the growth-differentiation balance hypothesis in Arabidopsis thaliana, Oecologia, vol.147, issue.4, pp.169-178, 2005.
DOI : 10.1016/S0176-1617(99)80020-0

J. E. Bassard, P. Ullmann, F. Bernier, and D. Werck-reichhart, Phenolamides: Bridging polyamines to the phenolic metabolism, Phytochemistry, vol.71, issue.16, pp.1808-1824, 2010.
DOI : 10.1016/j.phytochem.2010.08.003

F. A. Bazzaz, The Response of Natural Ecosystems to the Rising Global CO2 Levels, Annual Review of Ecology and Systematics, vol.21, issue.1, pp.167-196, 1990.
DOI : 10.1146/annurev.es.21.110190.001123

C. Bénard, Etude de l'impact de la nutrition azotée et des conditions de culture sur le contenu en polyphénols chez la tomate, 2009.

N. Benhamou, Elicitor-induced plant defence pathways, Trends in Plant Science, vol.1, issue.7, pp.233-240, 1996.
DOI : 10.1016/S1360-1385(96)86901-0

N. Benhamou, La résistance chez les plantes : Principes de la stratégie défensive et applications agronomiques, 2009.

N. Benhamou and K. Picard, Induced resistance??: a novel plant defense strategy against pathogens, Phytoprotection, vol.80, issue.3, pp.137-168, 1999.
DOI : 10.7202/706189ar

I. Berber, S. Ekin, P. Battal, H. Onlu, and M. E. Erez, Levels of Selected Trace Elements, Phytohormones, and Sugars in Pseudomonas-Infected Lycopersicum esculantum Mill Plants, Biological Trace Element Research, vol.50, issue.1, pp.98-109, 2010.
DOI : 10.1080/07352689991309397

S. Berger, M. Papadopoulos, U. Schreiber, W. Kaiser, and T. Roitsch, Complex regulation of gene expression, photosynthesis and sugar levels by pathogen infection in tomato, Physiologia Plantarum, vol.101, issue.4, pp.419-428, 2004.
DOI : 10.1006/pmpp.1995.1055

S. Berger, A. K. Sinha, and T. Roitsch, Plant physiology meets phytopathology: plant primary metabolism and plant pathogen interactions, Journal of Experimental Botany, vol.58, issue.15-16, pp.15-16, 2007.
DOI : 10.1093/jxb/erm298

M. J. Berlinger, Pests The tomato crop: A scientific basis for improvement, pp.391-432, 1986.

D. Blancard, Les maladies de la tomate: Identifier, connaître, maîtriser. Editions Quae, 2009.

R. P. Bodnaryk and P. Palaniswamy, Glucosinolate levels in cotyledons of mustard,Brassica juncea L. and rape,B. napus L. do not determine feeding rates of flea beetle,Phyllotreta cruciferae (Goeze), Journal of Chemical Ecology, vol.195, issue.10, pp.2735-2746, 1990.
DOI : 10.1007/BF00988082

K. B. Bonfig, U. Schreiber, A. Gabler, T. Roitsch, and S. Berger, Infection with virulent and avirulent P. syringae strains differentially affects photosynthesis and sink metabolism in Arabidopsis leaves, Planta, vol.136, issue.1, pp.1-12, 2006.
DOI : 10.1007/978-1-4020-3218-9_11

S. Bonny, L'intensification écologique de l'agriculture : voies et défis, Symposium "ISDA 2010 Innovation and Sustainable Development in Agriculture and Food, 2010.

A. J. Boughton, K. Hoover, and G. W. Felton, Impact of chemical elicitor applications on greenhouse tomato plants and population growth of the green peach aphid, Myzus persicae, Entomologia Experimentalis et Applicata, vol.257, issue.3, pp.175-188, 2006.
DOI : 10.1111/j.1365-294X.2004.02297.x

F. Bourgaud, A. Hehn, R. Larbat, S. Doerper, E. Gontier et al., Biosynthesis of coumarins in plants: a major pathway still to be unravelled for cytochrome P450 enzymes, Phytochemistry Reviews, vol.106, issue.2-3, pp.293-308, 2006.
DOI : 10.1515/bchm2.1971.352.1.78

URL : https://hal.archives-ouvertes.fr/hal-01738288

A. Bovy, E. Schijlen, and R. D. Hall, Metabolic engineering of flavonoids in tomato (Solanum lycopersicum): the potential for metabolomics, Metabolomics, vol.9, issue.3, pp.399-412, 2007.
DOI : 10.1093/oxfordjournals.pcp.a029312

P. D. Brown, J. G. Tokuhisa, M. Reichelt, and J. Gershenzon, Variation of glucosinolate accumulation among different organs and developmental stages of Arabidopsis thaliana, Phytochemistry, vol.62, issue.3, pp.471-481, 2003.
DOI : 10.1016/S0031-9422(02)00549-6

J. P. Bryant, F. S. Chapin, and D. R. Klein, Carbon/Nutrient Balance of Boreal Plants in Relation to Vertebrate Herbivory, Oikos, vol.40, issue.3, pp.357-368, 1983.
DOI : 10.2307/3544308

C. R. Buell, V. Joardar, M. Lindeberg, J. Selengut, I. T. Paulsen et al., The complete genome sequence of the Arabidopsis and tomato pathogen Pseudomonas syringae pv. tomato DC3000, Proceedings of the National Academy of Sciences of the United States of America, pp.10181-10186, 2003.
DOI : 10.1016/S1097-2765(00)80265-8

R. Cárdenas-navarro, S. Adamowicz, and P. Robin, Nitrate accumulation in plants: a role for water, Journal of Experimental Botany, vol.50, issue.334, pp.613-624, 1999.
DOI : 10.1093/jxb/50.334.613

O. Carpena, P. Zornoza, and J. Mataix, Incidence of P, Mn and B deficiencies on the levels of the whole and individual flavonoid groups in tomato leaves, Journal of Plant Nutrition, vol.3, issue.3, pp.1197-1208, 1982.
DOI : 10.1016/S0031-9422(00)83624-9

T. R. Cataldi, F. Lelario, and S. A. Bufo, Analysis of tomato glycoalkaloids by liquid chromatography coupled with electrospray ionization tandem mass spectrometry, Rapid Communications in Mass Spectrometry, vol.58, issue.21, pp.3103-3110, 2005.
DOI : 10.1002/rcm.2176

V. Catara, Pseudomonas corrugata: plant pathogen and/or biological resource?, Molecular Plant Pathology, vol.69, issue.9, pp.233-244, 2007.
DOI : 10.1111/j.1439-0434.1994.tb00007.x

S. Chaillou and T. Lamaze, Nutrition ammoniacale des plantes Assimilation de l'azote chez les plantes : Aspects physiologique, biochimique et moléculaire, pp.67-84, 1997.

A. Chang, M. H. Lim, S. W. Lee, E. J. Robb, and R. N. Nazar, Tomato phenylalanine ammonialyase gene family, highly redundant but strongly underutilized". The Journal of biological chemistry, pp.283-33591, 2008.

C. Clé, L. M. Hill, R. Niggeweg, C. R. Martin, Y. Guisez et al., Modulation of chlorogenic acid biosynthesis in Solanum lycopersicum; consequences for phenolic accumulation and UV-tolerance, Phytochemistry, issue.11, pp.69-2149, 2008.

D. B. Collinge, Cell wall appositions: the first line of defence, Journal of Experimental Botany, vol.36, issue.1, pp.351-352, 2009.
DOI : 10.1023/A:1005955119326

R. Corbaz, Maladies des plantes et leurs agents, Principes de phytopathologie et de lutte contre les maladies des plantes. PPUR presses polytechniques, pp.1-18, 1990.

J. M. Costa and E. Heuvelink, Introduction: the tomato crop and industry., Tomatoes, pp.1-20, 2005.
DOI : 10.1079/9780851993966.0001

W. J. Cox and H. M. Reisenauer, Growth and ion uptake by wheat supplied nitrogen as nitrate, or ammonium, or both, Plant and Soil, vol.1, issue.2, pp.363-380, 1973.
DOI : 10.1071/BI9480333

R. Croteau, T. M. Kutchan, and N. G. Lewis, Natural products (Secondary metabolites Biochemistry and molecular biology of plants, pp.1250-1318, 2000.

D. Jong, W. S. Eannetta, N. T. , D. Jong, D. M. Bodis et al., Candidate gene analysis of anthocyanin pigmentation loci in the Solanaceae, Theoretical and Applied Genetics, pp.423-432, 2004.
DOI : 10.1007/s00122-003-1455-1

C. M. De-oliveira, V. C. De-andrade, W. R. Maluf, I. P. Neiva, and G. M. Maciel, Resistance of tomato strains to the moth Tuta absoluta imparted by allelochemicals and trichome density, Ci??ncia e Agrotecnologia, vol.124, issue.36, pp.45-52, 2012.
DOI : 10.1590/S0100-204X2009001000008

J. T. De-resende, W. R. Maluf, M. V. Faria, A. Z. Pfann, and E. R. Do-nascimento, Acylsugars in tomato leaflets confer resistance to the South American tomato pinworm, Tuta absoluta Meyr, Scientia Agricola, vol.26, issue.1, pp.20-25, 2006.
DOI : 10.1023/A:1002963623325

V. Decognet, F. Ravetti, C. Martin, and P. C. Nicot, Improved leaf pruning reduces development of stem cankers caused by grey mould in greenhouse tomatoes, Agronomy for Sustainable Development, pp.465-472, 2010.
DOI : 10.1111/j.1364-3703.2007.00417.x

URL : https://hal.archives-ouvertes.fr/hal-00886532

D. C. Degenhardt, S. Refi-hind, J. W. Stratmann, and D. E. Lincoln, Systemin and jasmonic acid regulate constitutive and herbivore-induced systemic volatile emissions in tomato, Solanum lycopersicum, Phytochemistry, vol.71, issue.17-18, pp.2024-2037, 2010.
DOI : 10.1016/j.phytochem.2010.09.010

N. Desneux, M. G. Luna, T. Guillemaud, and A. Urbaneja, The invasive South American tomato pinworm, Tuta absoluta, continues to spread in Afro-Eurasia and beyond: the new threat to tomato world production, Journal of Pest Science, vol.96, issue.4, pp.403-408, 2011.
DOI : 10.1073/pnas.96.11.5944

N. Desneux, E. Wajnberg, K. A. Wyckhuys, G. Burgio, S. Arpaia et al., Biological invasion of European tomato crops by Tuta absoluta: ecology, geographic expansion and prospects for biological control, Journal of Pest Science, vol.36, issue.3, pp.197-215, 2010.
DOI : 10.1590/S0100-204X2002000500002

R. A. Dixon and N. L. Paiva, Stress-induced phenylpropanoid metabolism". The Plant Cell, pp.1085-1097, 1995.

T. Doré, D. Makowski, E. Malézieux, N. Munier-jolain, M. Tchamitchian et al., Facing up to the paradigm of ecological intensification in agronomy: Revisiting methods, concepts and knowledge, European Journal of Agronomy, vol.34, issue.4, pp.197-210, 2011.
DOI : 10.1016/j.eja.2011.02.006

N. Dudareva, F. Negre, D. A. Nagegowda, and I. Orlova, Plant Volatiles: Recent Advances and Future Perspectives, Critical Reviews in Plant Sciences, vol.2, issue.5, pp.417-440, 2006.
DOI : 10.1007/s10886-005-5285-2

N. Dudareva and E. Pichersky, Metabolic engineering of plant volatiles, Current Opinion in Biotechnology, vol.19, issue.2, pp.181-189, 2008.
DOI : 10.1016/j.copbio.2008.02.011

Y. Dumas, M. Dadomo, D. Lucca, G. Grolier, and P. , Effects of environmental factors and agricultural techniques on antioxidantcontent of tomatoes, Journal of the Science of Food and Agriculture, vol.277, issue.5, pp.369-382, 2003.
DOI : 10.17660/ActaHortic.1990.277.17

C. C. Ecole, M. C. Picanco, R. N. Guedes, and S. H. Brommonschenkel, Effect of cropping season and possible compounds involved in the resistance of Lycopersicon hirsutum f. typicum to Tuta absoluta (Meyrick) (Lep., Gelechiidae), Journal of Applied Entomology, vol.114, issue.4, pp.193-200, 2001.
DOI : 10.1016/0304-4211(84)90222-0

E. Oirdi, M. Abd-el-rahman, T. Rigano, L. Hadrami, A. Rodriguez et al., Manipulates the Antagonistic Effects between Immune Pathways to Promote Disease Development in Tomato, The Plant Cell, vol.23, issue.6, pp.2405-2421, 2011.
DOI : 10.1105/tpc.111.083394

M. Errebhi and G. E. Wilcox, Tomato growth and nutrient uptake pattern as influenced by nitrogen form ratio, Journal of Plant Nutrition, vol.111, issue.8, pp.1031-1043, 1990.
DOI : 10.1080/00103628509367673

P. J. Facchini, J. Hagel, and K. G. Zulak, Hydroxycinnamic acid amide metabolism: physiology and biochemistry, Canadian Journal of Botany, vol.108, issue.6, pp.577-589, 2002.
DOI : 10.1034/j.1399-3054.2000.108001101.x

J. W. Fahey, A. T. Zalcmann, and P. Talalay, The chemical diversity and distribution of glucosinolates and isothiocyanates among plants, Phytochemistry, vol.56, issue.1, pp.5-51, 2001.
DOI : 10.1016/S0031-9422(00)00316-2

J. D. Faure, C. Meyer, and M. Caboche, Assimilation du nitrate : nitrate et nitrite réductase Assimilation de l'azote chez les plantes, 1997.

A. Fleuriet and J. Macheix, Quinyl esters and glucose derivatives of hydroxycinnamic acids during growth and ripening of tomato fruit, Phytochemistry, vol.20, issue.4, pp.667-671, 1981.
DOI : 10.1016/0031-9422(81)85153-9

M. Friedman, Tomato Glycoalkaloids:?? Role in the Plant and in the Diet, Journal of Agricultural and Food Chemistry, vol.50, issue.21, pp.5751-5780, 2002.
DOI : 10.1021/jf020560c

M. Friedman, Analysis of biologically active compounds in potatoes (Solanum tuberosum), tomatoes (Lycopersicon esculentum), and jimson weed (Datura stramonium) seeds, Journal of Chromatography A, vol.1054, issue.1-2, pp.143-155, 2004.
DOI : 10.1016/j.chroma.2004.04.049

C. Fritz, C. Mueller, P. Matt, R. Feil, and M. Stitt, Impact of the C?N status on the amino acid profile in tobacco source leaves, Plant, Cell and Environment, vol.106, issue.11, pp.2055-2076, 2006.
DOI : 10.1105/tpc.10.3.359

C. Fritz, N. Palacios-rojas, R. Feil, and M. Stitt, Regulation of secondary metabolism by the carbon-nitrogen status in tobacco: nitrate inhibits large sectors of phenylpropanoid metabolism, The Plant Journal, vol.7, issue.4, pp.533-548, 2006.
DOI : 10.1016/0304-4165(78)90223-4

S. Gayler, T. E. Grams, W. Heller, D. Treutter, and E. Priesack, A Dynamical Model of Environmental Effects on Allocation to Carbon-based Secondary Compounds in Juvenile Trees, Annals of Botany, vol.101, issue.8, pp.1089-1098, 2008.
DOI : 10.1093/aob/mcm169

M. Geiger, V. Haake, F. Ludewig, U. Sonnewald, and M. Stitt, The nitrate and ammonium nitrate supply have a major influence on the response of photosynthesis, carbon metabolism, nitrogen metabolism and growth to elevated carbon dioxide in tobacco, Plant, Cell and Environment, vol.10, issue.10, pp.1177-1199, 1999.
DOI : 10.1071/PP9910137

I. Grechi, P. Vivin, G. Hilbert, S. Milin, T. Robert et al., Effect of light and nitrogen supply on internal C:N balance and control of root-to-shoot biomass allocation in grapevine, Environmental and Experimental Botany, vol.59, issue.2, pp.139-149, 2007.
DOI : 10.1016/j.envexpbot.2005.11.002

C. Grignon, J. B. Thibaud, and T. Lamaze, Assimilation de l'azote chez les plantes, 1997.

D. L. Gustine, R. T. Sherwood, B. G. Moyer, and F. L. Lukezic, Metabolites from pseudomonas corrugata elicit phytoalexin biosynthesis in white clover, Physiology and Biochemistry, vol.80, issue.12, pp.1427-1432, 1990.

J. G. Hamilton, A. R. Zangerl, E. H. Delucia, and M. R. Berenbaum, The carbon-nutrient balance hypothesis: its rise and fall, Ecology Letters, vol.5, issue.1, pp.86-95, 2001.
DOI : 10.1086/285868

K. E. Hammond-kosack and J. D. Jones, Resistance gene-dependent plant defense responses". The Plant Cell, pp.1773-1791, 1996.

R. J. Haynes and K. M. Goh, AMMONIUM AND NITRATE NUTRITION OF PLANTS, Biological Reviews, vol.15, issue.4, pp.465-510, 1978.
DOI : 10.1080/00380768.1969.10432789

M. Hehmann, R. Luka?in, H. Ekiert, and U. Matern, Furanocoumarin biosynthesis in Ammi majus L., European Journal of Biochemistry, vol.368, issue.5, pp.932-940, 2004.
DOI : 10.1016/0005-2744(72)90326-9

D. A. Herms and W. J. Mattson, The Dilemma of Plants: To Grow or Defend, The Quarterly Review of Biology, vol.67, issue.3, pp.283-335, 1992.
DOI : 10.1086/417659

E. Hoffland, M. Dicke, W. Van-tintelen, H. Dijkman, and M. L. Van-beusichem, Nitrogen availability and defense of tomato against two-spotted spider mite, Journal of Chemical Ecology, vol.26, issue.12, pp.2697-2711, 2000.
DOI : 10.1023/A:1026477423988

E. Hoffland, M. J. Jeger, and M. L. Van-beusichem, Effect of nitrogen supply rate on disease resistance in tomato depends on the pathogen, Plant and Soil, vol.218, issue.12, pp.239-247, 2000.

E. Hoffland, M. L. Van-beusichem, and M. J. Jeger, Nitrogen availability and susceptibility of tomato leaves to Botrytis cinerea, Plant and Soil, vol.210, issue.2, pp.263-272, 1999.
DOI : 10.1023/A:1004661913224

H. Magaña, R. Adamowicz, S. Pagès, and L. , Diel changes in nitrogen and carbon resource status and use for growth in young plants of tomato (Solanum lycopersicum), Annals of Botany, vol.16, issue.7, pp.1025-1037, 2009.
DOI : 10.1051/agro:19960201

D. M. Huber and I. A. Thompson, Nitrogen and plant disease Mineral nutrition and plant disease, pp.31-44, 2007.

M. H. Ibrahim, H. Z. Jaafar, A. Rahmat, and Z. A. Rahman, The Relationship between Phenolics and Flavonoids Production with Total Non Structural Carbohydrate and Photosynthetic Rate in Labisia pumila Benth. under High CO2 and Nitrogen Fertilization, Molecules, vol.16, issue.1, pp.162-174, 2011.
DOI : 10.3390/molecules16010162

K. Valimunzigha and C. , Etude du comportement physiologique et agronomique de la tomate (Solanum lycopersicum L.) en réponse à un stress hydrique précoce. Mémoire de thèse, 2006.

I. Kaplan, R. Halitschke, A. Kessler, S. Sardanelli, and R. F. Denno, CONSTITUTIVE AND INDUCED DEFENSES TO HERBIVORY IN ABOVE- AND BELOWGROUND PLANT TISSUES, Ecology, vol.108, issue.4, pp.392-406, 2008.
DOI : 10.1007/s004250050211

H. Keller, H. Hohlfeld, V. Wray, K. Hahlbrock, D. Scheel et al., Changes in the accumulation of soluble and cell wall-bound phenolics in elicitor-treated cell suspension cultures and fungus-infected leaves of Solanum tuberosum, Phytochemistry, vol.42, issue.2, pp.389-396, 1996.
DOI : 10.1016/0031-9422(95)00866-7

E. A. Keukens, T. De-vrije, C. Van-den-boom, P. De-waard, H. H. Plasman et al., Molecular basis of glycoalkaloid induced membrane disruption, Biochimica et Biophysica Acta (BBA) - Biomembranes, vol.1240, issue.2, pp.216-228, 1995.
DOI : 10.1016/0005-2736(95)00186-7

E. A. Keukens, M. E. Hop, and W. M. Jongen, Rapid high-performance liquid chromatographic method for the quantification of .alpha.-tomatine in tomato, Journal of Agricultural and Food Chemistry, vol.42, issue.11, pp.42-2475, 1994.
DOI : 10.1021/jf00047a020

J. Koricheva, S. Larsson, E. Haukioja, and M. Keinanen, Regulation of Woody Plant Secondary Metabolism by Resource Availability: Hypothesis Testing by Means of Meta-Analysis, Oikos, vol.83, issue.2, pp.212-226, 1998.
DOI : 10.2307/3546833

N. Kozukue, J. Han, K. Lee, and M. Friedman, Dehydrotomatine and ??-Tomatine Content in Tomato Fruits and Vegetative Plant Tissues, Journal of Agricultural and Food Chemistry, vol.52, issue.7, pp.2079-2083, 2004.
DOI : 10.1021/jf0306845

R. Larbat, L. Bot, J. Bourgaud, F. Robin, C. Adamowicz et al., Organ-specific responses of tomato growth and phenolic metabolism to nitrate limitation, Plant Biology, vol.77, issue.5, pp.760-769, 2012.
DOI : 10.2307/2265674

R. Larbat, K. M. Olsen, R. Slimestad, T. Løvdal, C. Bénard et al., Influence of repeated short-term nitrogen limitations on leaf phenolics metabolism in tomato, Phytochemistry, vol.77, pp.119-147, 2012.
DOI : 10.1016/j.phytochem.2012.02.004

L. Bot, J. Bénard, C. Robin, C. Bourgaud, F. Adamowicz et al., The ???trade-off??? between synthesis of primary and secondary compounds in young tomato leaves is altered by nitrate nutrition: experimental evidence and model consistency, Journal of Experimental Botany, vol.82, issue.15, pp.4301-4314, 2009.
DOI : 10.1002/jsfa.1267

F. Lecompte, M. A. Abro, and P. C. Nicot, Contrasted responses of Botrytis cinerea isolates developing on tomato plants grown under different nitrogen nutrition regimes, Plant Pathology, vol.55, issue.5, pp.891-899, 2010.
DOI : 10.1111/j.1365-3059.2010.02320.x

E. Lee and K. Paek, Effect of nitrogen source on biomass and bioactive compound production in submerged cultures of Eleutherococcus koreanum nakai adventitious roots, Biotechnology Progress, vol.30, issue.2, pp.508-514, 2012.
DOI : 10.5010/JPB.2003.30.3.301

G. L. Leite, M. Picanço, A. A. Azevedo, D. J. Silva, and M. R. Gusmão, Intensidade de ataque de Frankliniella schulzei e Myzus persicae em três introduções de Lycopersicon peruvianum, revista da Universidade Rural, Série Ciência da Vida, pp.27-35, 1997.

G. L. Leite, M. Picanço, G. N. Jham, and F. Marquini, Intensity of Tuta absoluta (Meyrick, 1917) (Lepidoptera: Gelechiidae) and Liriomyza spp, Diptera: Agromyzidae) attacks on Lycopersicum esculentum Mill. leaves". Ciencia E Agrotecnologia, pp.42-48, 2004.

G. L. Leite, M. Picanco, J. C. Zanuncio, M. D. Moreira, and G. N. Jham, Hosting Capacity of Horticultural Plants for Insect Pests in Brazil, Chilean journal of agricultural research, vol.71, issue.3, pp.383-389, 2011.
DOI : 10.4067/S0718-58392011000300006

D. K. Letourneau, L. E. Drinkwater, and C. Shennan, Effects of soil management on crop nitrogen and insect damage in organic vs. conventional tomato fields, Agriculture, Ecosystems & Environment, vol.57, issue.2-3, pp.2-3, 1996.
DOI : 10.1016/0167-8809(96)01027-4

J. Li, J. M. Zhou, and Z. Q. Duan, Effects of elevated CO2 concentration on growth and water usage of tomato seedlings under different ammonium/nitrate ratios, Journal of Environmental Sciences, vol.19, issue.9, pp.1100-1107, 2007.
DOI : 10.1016/S1001-0742(07)60179-X

J. Y. Li, X. H. Liu, Q. S. Cai, H. Gu, S. S. Zhang et al., on Growth, Carbon Assimilation, Photosynthate Accumulation and Related Enzymes in Rice Leaves during Sink-Source Transition, Journal of Integrative Plant Biology, vol.161, issue.6, pp.723-732, 2008.
DOI : 10.1093/oxfordjournals.pcp.a029608

W. E. Loomis, Growth-differentiation balance vs. carbohydrate-nitrogen ratio, Proceedings of the American Society of Horticultural Sciences, pp.240-245, 1932.

M. P. López-gresa, F. Maltese, J. M. Bellés, V. Conejero, H. K. Kim et al., Metabolic response of tomato leaves upon different plant??????pathogen interactions, Phytochemical Analysis, vol.20, issue.1, pp.89-94, 2010.
DOI : 10.1007/978-3-642-74164-7_12

M. P. López-gresa, C. Torres, L. Campos, P. Lisón, I. Rodrigo et al., Identification of defence metabolites in tomato plants infected by the bacterial pathogen Pseudomonas syringae, Environmental and Experimental Botany, vol.74, pp.216-228, 2011.
DOI : 10.1016/j.envexpbot.2011.06.003

J. Lopez, N. Tremblay, W. Voogt, S. Dubé, and A. Gosselin, Effects of varying sulphate concentrations on growth, physiology and yield of the greenhouse tomato, Scientia Horticulturae, vol.67, issue.3-4, pp.207-217, 1996.
DOI : 10.1016/S0304-4238(96)00948-X

T. Løvdal and C. Lillo, Reference gene selection for quantitative real-time PCR normalization in tomato subjected to nitrogen, cold, and light stress, Analytical Biochemistry, vol.387, issue.2, pp.238-242, 2009.
DOI : 10.1016/j.ab.2009.01.024

T. Løvdal, K. M. Olsen, R. Slimestad, M. Lillo, and C. , Synergetic effects of nitrogen depletion, temperature, and light on the content of phenolic compounds and gene expression in leaves of tomato, Phytochemistry, vol.71, issue.5-6, pp.605-613, 2010.
DOI : 10.1016/j.phytochem.2009.12.014

J. J. Macheix, A. Fleuriet, and C. Jay-allemand, Les composés phénoliques des végétaux: un exemple de métabolites secondaires d'importance économique, Presses Polytechniques et Universitaires Romandes, 2005.

E. A. Maher, N. J. Bate, W. Ni, Y. Elkind, R. A. Dixon et al., Increased disease susceptibility of transgenic tobacco plants with suppressed levels of preformed phenylpropanoid products., Proceedings of the National Academy of Sciences, pp.91-7802, 1994.
DOI : 10.1073/pnas.91.16.7802

J. Malamy, J. P. Carr, D. F. Klessig, and I. Raskin, Salicylic Acid: A Likely Endogenous Signal in the Resistance Response of Tobacco to Viral Infection, Science, vol.250, issue.4983, pp.250-1002, 1990.
DOI : 10.1126/science.250.4983.1002

W. R. Maluf, G. M. Maciel, L. A. Gomes, M. D. Cardoso, L. D. Goncalves et al., Broad-Spectrum Arthropod Resistance in Hybrids between High-and Low-Acylsugar Tomato Lines". Crop science, pp.439-450, 2010.

H. Marschner, Relationship between mineral nutrition and plant diseases and pests Mineral nutrition of higher plants (seconde édition), pp.436-460, 1995.

A. Matros, S. Amme, B. Kettig, G. H. Buck-sorlin, U. Sonnewald et al., Growth at elevated CO2 concentrations leads to modified profiles of secondary metabolites in tobacco cv. SamsunNN and to increased resistance against infection with potato virus Y, Plant, Cell and Environment, vol.134, issue.1, pp.126-137, 2006.
DOI : 10.1146/annurev.phyto.27.1.143

J. W. Mattson, R. Julkunen-tiitto, and D. A. Herms, CO 2 enrichment and carbon partitioning to phenolics: do plant responses accord better with the protein competition or the growth differentiation balance models?" Oikos, pp.337-347, 2005.

D. Mckey, Adaptive Patterns in Alkaloid Physiology, The American Naturalist, vol.108, issue.961, pp.305-320, 1974.
DOI : 10.1086/282909

I. Mehrer and H. Mohr, Ammonium toxicity: description of the syndrome in Sinapis alba and the search for its causation, Physiologia Plantarum, vol.41, issue.4, pp.545-554, 1989.
DOI : 10.1016/0003-2697(83)90117-3

F. Mert-türk, Phytoalexins: Defence or just a response to stress ?, Journal of Cell and Molecular Biology, vol.1, pp.1-6, 2002.

S. Meyer, C. Reeb, and R. Bosdeveix, Botanique, biologie et physiologie végétales, 2004.

C. A. Mihaliak and D. E. Lincoln, Growth pattern and carbon allocation to volatile leaf terpenes under nitrogen-limiting conditions in Heterotheca subaxillaris (Asteraceae), Oecologia, vol.62, issue.3, pp.423-426, 1985.
DOI : 10.1007/BF00378309

S. E. Milner, N. P. Brunton, P. W. Jones, N. M. O-'brien, S. G. Collins et al., Bioactivities of Glycoalkaloids and Their Aglycones from Solanum Species, Journal of Agricultural and Food Chemistry, vol.59, issue.8, pp.59-3454, 2011.
DOI : 10.1021/jf200439q

K. Mittelstraß, D. Treutter, M. Pleßl, W. Heller, E. F. Elstner et al., Modification of Primary and Secondary Metabolism of Potato Plants by Nitrogen Application Differentially Affects Resistance to Phytophthora infestans and Alternaria solani, Plant Biology, vol.70, issue.5, pp.653-661, 2006.
DOI : 10.1105/tpc.7.11.1787

S. Moco, R. J. Bino, O. Vorst, H. A. Verhoeven, J. De-groot et al., A Liquid Chromatography-Mass Spectrometry-Based Metabolome Database for Tomato, PLANT PHYSIOLOGY, vol.141, issue.4, pp.1205-1218, 2006.
DOI : 10.1104/pp.106.078428

O. Mollà, H. Montón, P. Vanaclocha, F. Bieitia, and A. Urbaneja, Predation by the mirids Nesidiocoris tenuis and Macrolophus pygmaeus on the tomato borer Tuta absoluta, 2009.

I. Group, Integrated Control in Protected Crops, Mediterranean Climate, Crète, pp.209-214

M. Montanari, E. Degl-'innocenti, R. Maggini, S. Pacifici, A. Pardossi et al., Effect of nitrate fertilization and saline stress on the contents of active constituents of Echinacea angustifolia DC". Food Chemistry, pp.1461-1466, 2008.

J. F. Morot-gaudry, Métabolisme secondaire : quelques aspects, in: Biologie végétale Croissance et développement, pp.217-227, 2012.

C. E. Morris, D. C. Sands, B. A. Vinatzer, C. Glaux, C. Guilbaud et al., The life history of the plant pathogen Pseudomonas syringae is linked to the water cycle, The ISME Journal, vol.146, issue.3, pp.321-334, 2008.
DOI : 10.1099/00221287-146-10-2385

I. M. Morrison and D. Stewart, Determination of lignin in the presence of ester-bound substituted cinnamic acids by a modified acetyl bromide procedure, Journal of the Science of Food and Agriculture, vol.39, issue.2, pp.151-157, 1995.
DOI : 10.1007/978-1-349-15203-2

S. Naika, J. Van-lidt-de-jeude, M. De-goffau, and B. Van-dam, La culture de la tomate production, transformation et commercialisation Agromisa Foundation, 2005.

R. Niggeweg, A. Michael, and C. Martin, Engineering plants with increased levels of the antioxidant chlorogenic acid, Nature Biotechnology, vol.10, issue.6, pp.746-754, 2004.
DOI : 10.1105/tpc.10.11.1801

L. H. Nugroho, A. M. Peltenburg-looman, H. De-vos, M. C. Verberne, and R. Verpoorte, Nicotine and related alkaloids accumulation in constitutive salicylic acid producing tobacco plants, Plant Science, vol.162, issue.4, pp.575-581, 2002.
DOI : 10.1016/S0168-9452(01)00596-9

T. R. Nuringtyas, Y. H. Choi, R. Verpoorte, P. G. Klinkhamer, and K. A. Leiss, Differential tissue distribution of metabolites in Jacobaea vulgaris, Jacobaea aquatica and their crosses, Phytochemistry, vol.78, pp.89-97, 2012.
DOI : 10.1016/j.phytochem.2012.03.011

A. E. Osbourn, Preformed antimicrobial compounds and plant defense against fungal attack". The Plant Cell, pp.1821-1831, 1996.

F. Papy and I. Goldringer, Cultiver la biodiversité". Courrier de l'environnement de l'INRA, pp.55-62, 2011.

S. Park, W. Y. Jeong, J. H. Lee, Y. Kim, S. W. Jeong et al., Determination of polyphenol levels variation in Capsicum annuum L. cv. Chelsea (yellow bell pepper) infected by anthracnose (Colletotrichum gloeosporioides) using liquid chromatography???tandem mass spectrometry, Food Chemistry, vol.130, issue.4, pp.981-985, 2012.
DOI : 10.1016/j.foodchem.2011.08.026

M. M. Peet and G. W. Welles, Greenhouse tomato production., Tomatoes, pp.257-305, 2005.
DOI : 10.1079/9780851993966.0257

J. M. Peñalosa, M. J. Sarro, E. Revilla, R. Carpena, and C. Cadahía, Influence of phosphorus supply on tomato plant nutrition, Journal of Plant Nutrition, vol.60, issue.5, pp.647-657, 1989.
DOI : 10.1007/BF01372881

J. Peñuelas and M. Estiarte, Can elevated CO2 affect secondary metabolism and ecosystem function?, Trends in Ecology & Evolution, vol.13, issue.1, pp.20-24, 1998.
DOI : 10.1016/S0169-5347(97)01235-4

J. Peñuelas, M. Estiarte, and J. Llusià, Carbon-based secondary compounds at elevated CO 2, Photosynthetica, vol.33, issue.2, pp.313-316, 1997.
DOI : 10.1023/A:1022120431279

G. V. Pereira, W. R. Maluf, L. D. Goncalves, D. R. Do-nascimento, L. A. Gomes et al., Sele????o para alto teor de acila????cares em gen??tipos de tomateiro e sua rela????o com a resist??ncia ao ??caro vermelho (Tetranychus evansi) e ?? tra??a (Tuta absoluta), Ci??ncia e Agrotecnologia, vol.114, issue.3, pp.996-1004, 2008.
DOI : 10.1023/A:1012067505361

C. M. Pieterse, A. Leon-reyes, S. Van-der-ent, and S. C. Van-wees, Networking by small-molecule hormones in plant immunity, Nature Chemical Biology, vol.24, issue.5, pp.308-316, 2009.
DOI : 10.1016/j.copbio.2006.01.009

D. S. Pitchay, Impact of applied nitrogen concentration on growth of elatior Begonia and New Guinea Impatiens and susceptibility of Begonia to Botrytis cinerea, Journal of the American Society for Horticultural Science, vol.132, pp.193-201, 2007.

B. W. Poovaiah, A. S. Reddy, and L. Feldman, Calcium and Signal Transduction in Plants, Critical Reviews in Plant Sciences, vol.84, issue.3, pp.185-211, 1993.
DOI : 10.1073/pnas.84.11.3580

T. Quidde, A. E. Osbourn, and P. Tudzynski, Detoxification of ?-tomatine by Botrytis cinerea". Physiological and Molecular Plant Pathology, pp.151-165, 1998.

R. Houma, A. Chérif, M. Boubaker, and A. , Effect of nitrogen fertilizing, green pruning and fungicide treatments on Botrytis bunch rot of grapes, Journal of Plant Pathology, vol.80, pp.115-124, 1998.

I. Radoj?i?-redovnikovi?, T. Gliveti?, K. Delonga, and J. Vorkapi?-fura?, Glucosinolates and their potential role in plant, Periodicum Biologorum, vol.110, issue.4, pp.297-309, 2008.

A. Raffo, C. Leonardi, V. Fogliano, P. Ambrosino, M. Salucci et al., Nutritional value of cherry tomatoes (Lycopersicum esculentum Cv Naomi F1) harvested at different ripening stages, Journal of Agricultural and Food Chemistry, issue.22, pp.50-6550, 2002.

T. Raghava, P. Ravikumar, R. Hegde, and A. Kush, Spatial and temporal volatile organic compound response of select tomato cultivars to herbivory and mechanical injury, Plant Science, vol.179, issue.5, pp.520-526, 2010.
DOI : 10.1016/j.plantsci.2010.07.020

A. H. Rasmy, The biology of the two-spotted spider mite Tetranychus urticae as affected by resistant solanaceous plants, Agriculture, Ecosystems & Environment, vol.13, issue.3-4, pp.3-4, 1985.
DOI : 10.1016/0167-8809(85)90020-9

M. Reisdorf-cren, La plante et son environnement biotique, in: Biologie végétale : croissance et développement, pp.187-216, 2012.

G. Ricochon, C. Paris, M. Girardin, and L. Muniglia, Highly sensitive, quick and simple quantification method for mono and disaccharides in aqueous media using liquid chromatography???atmospheric pressure chemical ionization???mass spectrometry (LC???APCI???MS), Journal of Chromatography B, vol.879, issue.19, pp.1529-1536, 2011.
DOI : 10.1016/j.jchromb.2011.03.044

M. R. Roberts and N. D. Paul, Seduced by the dark side: integrating molecular and ecological perspectives on the influence of light on plant defence against pests and pathogens, New Phytologist, vol.17, issue.4, pp.677-699, 2006.
DOI : 10.1080/07060669509500714

M. Royer, R. Larbat, L. Bot, J. Adamowicz, S. Robin et al., Is the C:N ratio a reliable indicator of C allocation to primary and defence-related metabolisms in tomato?, Phytochemistry, vol.88, 2013.
DOI : 10.1016/j.phytochem.2012.12.003

URL : https://hal.archives-ouvertes.fr/hal-01474765

J. Ruan, R. Haerdter, and J. Gerendás, Impact of nitrogen supply on carbon/nitrogen allocation: a case study on amino acids and catechins in green tea [Camellia sinensis (L.) O. Kuntze] plants*, Plant Biology, vol.126, issue.5, pp.724-734, 2010.
DOI : 10.1080/00380768.1995.10417026

T. W. Rufty, D. M. Jackson, R. F. Severson, J. J. Lam, and M. E. Snook, Alterations in growth and chemical constituents of tobacco in response to carbon dioxide enrichment, Journal of Agricultural and Food Chemistry, vol.37, issue.2, pp.552-555, 1989.
DOI : 10.1021/jf00086a060

J. M. Ruiz, G. Bretones, M. Baghour, L. Ragala, A. Belakbir et al., Relationship between boron and phenolic metabolism in tobacco leaves, Phytochemistry, vol.48, issue.2, pp.269-272, 1998.
DOI : 10.1016/S0031-9422(97)01132-1

J. M. Ruiz, R. M. Rivero, I. Lopez-cantarero, and L. Romero, Role of Ca 2+ in the metabolism of phenolic compounds in tobacco leaves (Nicotiana tabacum L.)". Plant Growth Regulation, pp.173-177, 2003.

J. A. Ryals, U. H. Neuenschwander, M. G. Willits, A. Molina, H. Y. Steiner et al., Systemic Acquired Resistance, THE PLANT CELL ONLINE, vol.8, issue.10, pp.1809-1819, 1996.
DOI : 10.1105/tpc.8.10.1809

E. Sánchez-rodríguez, J. M. Ruiz, F. Ferreres, and D. A. Moreno, Phenolic profiles of cherry tomatoes as influenced by hydric stress and rootstock technique, Food Chemistry, vol.134, issue.2, pp.775-782, 2012.
DOI : 10.1016/j.foodchem.2012.02.180

S. Sato, S. Tabata, H. Hirakawa, E. Asamizu, K. Shirasawa et al., The tomato genome sequence provides insights into fleshy fruit evolution, Nature, vol.63, issue.7400, pp.485-635, 2012.
DOI : 10.1007/s00239-005-0306-x

M. Schädler, M. Roeder, R. Brandl, and D. Matthies, , nutrient availability and plant species on a generalist invertebrate herbivore, Global Change Biology, vol.26, issue.5, pp.1005-1015, 2007.
DOI : 10.1046/j.1365-2486.2002.00497.x

D. Scheel, Resistance response physiology and signal transduction, Current Opinion in Plant Biology, vol.1, issue.4, pp.305-310, 1998.
DOI : 10.1016/1369-5266(88)80051-7

W. R. Scheible, A. Gonzalez-fontes, M. Lauerer, B. Muller-rober, M. Caboche et al., Nitrate Acts as a Signal to Induce Organic Acid Metabolism and Repress Starch Metabolism in Tobacco, THE PLANT CELL ONLINE, vol.9, issue.5, pp.783-798, 1997.
DOI : 10.1105/tpc.9.5.783

G. L. Shadle, S. V. Wesley, K. L. Korth, F. Chen, C. Lamb et al., Phenylpropanoid compounds and disease resistance in transgenic tobacco with altered expression of l -phenylalanine ammonia-lyase, Phytochemistry, vol.64, issue.1, pp.153-161, 2003.
DOI : 10.1016/S0031-9422(03)00151-1

R. Slimestad and M. J. Verheul, Seasonal Variations in the Level of Plant Constituents in Greenhouse Production of Cherry Tomatoes, Journal of Agricultural and Food Chemistry, vol.53, issue.8, pp.3114-3119, 2005.
DOI : 10.1021/jf047864e

S. Smole? and W. Sady, The effect of various nitrogen fertilization and foliar nutrition regimes on the concentrations of sugars, carotenoids and phenolic compounds in carrot (Daucus carota L.), Scientia Horticulturae, vol.120, issue.3, pp.315-324, 2009.
DOI : 10.1016/j.scienta.2008.11.029

P. S. Solomon, K. Tan, and R. P. Oliver, The nutrient supply of pathogenic fungi; a fertile field for study, Molecular Plant Pathology, vol.216, issue.3, pp.203-210, 2003.
DOI : 10.1006/pmpp.2000.0286

N. Stamp, Out Of The Quagmire Of Plant Defense Hypotheses, The Quarterly Review of Biology, vol.78, issue.1, pp.23-55, 2003.
DOI : 10.1086/367580

N. Stamp, Can the growth-differentiation balance hypothesis be tested rigorously?, Oikos, vol.68, issue.2, pp.439-448, 2004.
DOI : 10.1080/11956860.1997.11682411

M. Stitt, Nitrate regulation of metabolism and growth, Current Opinion in Plant Biology, vol.2, issue.3, pp.178-186, 1999.
DOI : 10.1016/S1369-5266(99)80033-8

M. Stitt and A. Krapp, The interaction between elevated carbon dioxide and nitrogen nutrition: the physiological and molecular background, Plant, Cell and Environment, vol.23, issue.6, pp.583-621, 1999.
DOI : 10.1071/PP9960045

URL : https://hal.archives-ouvertes.fr/hal-01608545

M. J. Stout, R. A. Brovont, and S. S. Duffey, Effect of nitrogen availability on expression of constitutive and inducible chemical defenses in tomato, Lycopersicon esculentum, Journal of Chemical Ecology, vol.24, issue.6, pp.945-963, 1998.
DOI : 10.1023/A:1022350100718

J. S. Thaler, Induced Resistance in Agricultural Crops: Effects of Jasmonic Acid on Herbivory and Yield in Tomato Plants, Environmental Entomology, vol.28, issue.1, pp.30-37, 1999.
DOI : 10.1093/jee/82.3.935

R. Tischner, W. Kaiser, H. Bothe, S. J. Ferguson, and N. , Nitrate Assimilation in Plants, Biology of the Nitrogen Cycle, 2007.
DOI : 10.1016/B978-044452857-5.50019-9

R. K. Toor, G. P. Savage, and A. Heeb, Influence of different types of fertilisers on the major antioxidant components of tomatoes, Journal of Food Composition and Analysis, vol.19, issue.1, pp.20-27, 2006.
DOI : 10.1016/j.jfca.2005.03.003

M. S. Traugott and N. Stamp, Effects of chlorogenic acid- and tomatine-fed caterpillars on performance of an insect predator, Oecologia, vol.109, issue.2, pp.265-272, 1997.
DOI : 10.1007/s004420050082

D. Treutter, Significance of flavonoids in plant resistance: a review, Environmental Chemistry Letters, vol.12, issue.Suppl, pp.147-157, 2006.
DOI : 10.1094/MPMI.2002.15.12.1204

E. Urbanczyk-wochniak and A. R. Fernie, Metabolic profiling reveals altered nitrogen nutrient regimes have diverse effects on the metabolism of hydroponically-grown tomato (Solanum lycopersicum) plants, Journal of Experimental Botany, vol.56, issue.410, pp.309-321, 2005.
DOI : 10.1104/pp.127.1.345

P. Van-baarlen, L. Legendre, and J. A. Van-kan, Plant defence compounds against Botrytis infection Botrytis: biology, pathology and control, pp.143-161, 2004.

J. Van-de-vooren, G. W. Welles, and G. Hayman, Glasshouse crop production The tomato crop : A scientific basis for improvement, pp.581-624, 1986.

H. D. Van-etten, J. W. Mansfield, J. A. Bailey, and E. E. Farmer, Two Classes of Plant Antibiotics: Phytoalexins versus "Phytoanticipins", THE PLANT CELL ONLINE, vol.6, issue.9, pp.1191-1192, 1994.
DOI : 10.1105/tpc.6.9.1191

G. Vercambre and S. Adamowicz, Dosage de l'ion nitrate en solution nutritive et en pr??sence de poly??thyl??ne glycol par spectrom??trie UV, Agronomie, vol.16, issue.2, pp.73-87, 1996.
DOI : 10.1051/agro:19960201

URL : http://www.agronomy-journal.org/articles/agro/pdf/1996/02/Agronomie_0249-5627_1996_16_2_ART0001.pdf

R. Verpoorte and J. Memelink, Engineering secondary metabolite production in plants, Current Opinion in Biotechnology, vol.13, issue.2, pp.181-187, 2002.
DOI : 10.1016/S0958-1669(02)00308-7

R. Verpoorte, R. Van-der-heijden, and J. Schripsema, Plant Cell Biotechnology for the Production of Alkaloids: Present Status and Prospects, Journal of Natural Products, vol.56, issue.2, pp.186-207, 1993.
DOI : 10.1021/np50092a003

E. A. Vidal and R. A. Gutiérrez, A systems view of nitrogen nutrient and metabolite responses in Arabidopsis, Current Opinion in Plant Biology, vol.11, issue.5, pp.521-529, 2008.
DOI : 10.1016/j.pbi.2008.07.003

J. C. Watterson, The tomato crop: A scientific basis for improvement, Disease, pp.443-484, 1986.

R. T. Wilkens, J. M. Spoerke, and N. E. Stamp, Differential Responses of Growth and Two Soluble Phenolics of Tomato to Resource Availability, Ecology, vol.77, issue.1, pp.247-258, 1996.
DOI : 10.2307/2265674

S. Yahara, N. Uda, and T. Nohara, Lycoperosides A-C, three stereoisomeric 23-acetoxyspirosolan-3??-ol ??-lycotetraosides from Lycopersicon esculentum, Phytochemistry, vol.42, issue.1, pp.169-172, 1996.
DOI : 10.1016/0031-9422(95)00854-3

U. Yermiyahu, I. Shamai, R. Peleg, N. Dudai, and D. Shtienberg, Reduction of Botrytis cinerea sporulation in sweet basil by altering the concentrations of nitrogen and calcium in the irrigation solution, Plant Pathology, vol.81, issue.4, pp.544-552, 2006.
DOI : 10.1007/BF01974242

A. M. Zobel and S. A. Brown, Seasonal changes of furanocoumarin concentrations in leaves ofHeracleum lanatum, Journal of Chemical Ecology, vol.16, issue.5, pp.1623-1634, 1990.
DOI : 10.1016/0165-1110(76)90012-9

P. Zornoza, J. Sanz, and O. Carpena, Analysis of the interaction between light and the type of nitrogen nutrition in tomato plants, Suelo y Planta, vol.1, issue.3, pp.389-398, 1991.

L. S. Adamowicz and J. Bor, preserves the proportionate relation linking long-term organic-nitrogen accumulation to intercepted radiation, New Phytologist, vol.59, issue.3, pp.663-672, 2008.
DOI : 10.2135/cropsci1994.0011183X003400030022x

S. Adamowicz, L. J. Bor, R. Huanosro-magaiia, and J. Fabre, Totomatix: a novel automatic set-up to control diurnal, diel and long-term plant nitrate nutrition, Annals of Botany, vol.16, issue.1, 2012.
DOI : 10.1051/agro:19960201

I. T. Baldwin, M. J. Karb, and T. E. Ohnmeiss, Allocation of 15N from Nitrate to Nicotine: Production and Turnover of a Damage-Induced Mobile Defense, Ecology, vol.75, issue.6, pp.1703-1713, 1994.
DOI : 10.2307/1939630

S. Bazor, J. Mikola, C. Nguyen, and C. Robin, Oefoliation-induced changes 1n carbon allocation and root soluble carbon concentration in field-grown Loüum perenne plants: d o they affect carbon availability, microbes and animal trophic groups in soil ?, Funct. Ecol, vol.19, pp.886-896, 2005.

F. A. Bazzaz, The Response of Natural Ecosystems to the Rising Global CO2 Levels, Annual Review of Ecology and Systematics, vol.21, issue.1, pp.167-196, 1990.
DOI : 10.1146/annurev.es.21.110190.001123

C. Bénard, H. Gautier, . F. Bourgaud, D. Grasselly, B. Navez et al., Effects of low nitrogen supply on tomate (Solanwn lycopersicwn) fruit yield and quality with special emphasis on sugars, acids, ascorbate, carotenoids, and phenolic compounds, J. Agric, pp.41-53, 2009.

R. P. Bodnaryk and P. Palaniswamy, Glucosinolate levels in cotyledons of mustard,Brassica juncea L. and rape,B. napus L. do not determine feeding rates of flea beetle,Phyllotreta cruciferae (Goeze), Journal of Chemical Ecology, vol.195, issue.10, pp.2735-2746, 1990.
DOI : 10.1007/BF00988082

J. P. Bryant, F. S. Cha-pin, O. R. Klein, R. Navarro, S. Adamowicz et al., Carbon/Nutrient Balance of Boreal Plants in Relation to Vertebrate Herbivory, Oikos, vol.40, issue.3, pp.357-368, 1983.
DOI : 10.2307/3544308

T. R. Cataldi, F. Lelario, . Bufo, and . Sa, Analysis of tomato glycoalkaloids by liquid chromatography coupled with electrospray ionization tandem mass spectrometry, Rapid Communications in Mass Spectrometry, vol.58, issue.21, pp.3103-3110, 2005.
DOI : 10.1002/rcm.2176

. W. Cox and H. M. Reisenauer, Growth and ion u ptake by whear supplied nitrogen as nitr are, or ammonium, or both, Plant Soi!, vol.38, pp.363-380, 1973.

R. Croreau, T. M. Kutchan, and N. G. Lewis, Natural products (secondary metabolites), Biochemistry and Molecular Biology of Plants. American Society of Plant Nutrition, pp.1250-1318, 2000.

A. Fleuriet and J. J. Macheix, Effect of anaerobie condi tions on the phenolic com pounds of 'Cheny' tomatoes, Lycopersicum esculentum var, Cerasiforme. Physiol. Veg, vol.4, pp.407-414, 1976.

M. Friedman, Tomato Glycoalkaloids:?? Role in the Plant and in the Diet, Journal of Agricultural and Food Chemistry, vol.50, issue.21, pp.5751-5780, 2002.
DOI : 10.1021/jf020560c

. C. Fritz, N. Palacios-rojas, . R. Feil, and M. Stitr, Regulation of secondary metabolism by the carbon-nitrogen status in tobacco: nitrate inhibits large sectors of phenylpropanoid metabolism, The Plant Journal, vol.7, issue.4, pp.533-548, 2006.
DOI : 10.1016/0304-4165(78)90223-4

M. Geiger, V. Haake, F. Ludewig, U. Sonnewald, and M. Stitr, The nitrate and ammonium nitrate supply have a major influence on the response of photosynthesis, carbon metabolism, nitrogen metabolism and growth to elevated carbon dioxide in tobacco, Plant, Cell and Environment, vol.10, issue.10, pp.1177-1199, 1999.
DOI : 10.1071/PP9910137

C. Graham, Nonstructural carbohydrate and prunasin composition of peach seedlings fertilized with different nitrogen sources and aluminum, Scientia Horticulturae, vol.94, issue.1-2, pp.21-32, 2002.
DOI : 10.1016/S0304-4238(01)00345-4

. Grechl-1, P. Vivin, . G. Hilbert, S. Mil-in, . T. Robert et al., Effect of light and nitrogen supply on internai C:N balance and control of root-to-shoot biomass a llocation in grapevine, pp.139-149, 2007.

R. J. Haynes and K. M. Goh, AMMONIUM AND NITRATE NUTRITION OF PLANTS, Biological Reviews, vol.15, issue.4, pp.465-470, 1978.
DOI : 10.1080/00380768.1969.10432789

D. A. Herms, Effects of fertilization on insect resistance of woody orna mental plants: reassessing an entrenched paradigm, Environ. Entomol, vol.3, issue.1, pp.923-933, 2002.

D. A. Herms and W. J. Mattson, The Dilemma of Plants: To Grow or Defend, The Quarterly Review of Biology, vol.67, issue.3, p.283, 1992.
DOI : 10.1086/417659

E. Hoffland, M. Dicke, W. Van-tintelen, H. Dijkman, and M. L. Van-beusichem, Nitrogen availability and defense of tomato against rwo-spotted spider mite, J, 2000.

E. Hoffland, M. L. Van-beusichem, and M. J. Jeger, Nitrogen availability and susceptibility of tomate leaves to Botrytis cinerea, Plant and Soil, vol.210, issue.2, pp.263-272, 1999.
DOI : 10.1023/A:1004661913224

F. Horchanl, R. Hajrl, and S. Aschi-smiti, Effect of ammonium or nitrate nutrition on photosynthesis, growth, and nitrogen assimilation in tomato plants, Journal of Plant Nutrition and Soil Science, vol.70, issue.4, pp.610-617
DOI : 10.1002/jpln.201000055

H. Magaiia, R. Adamowicz, S. Pagès, and L. , Die! changes in nitrogen and carbon resource status and use for growth in young plants of romato (Soltmum lycopersicum), Ann. Bot, vol.103, pp.1025-1037, 2009.

M. H. Ibrahim, H. Z. Jaafa-r, A. Rahmar, and Z. A. Rahman, The Relationship between Phenolics and Flavonoids Production with Total Non Structural Carbohydrate and Photosynthetic Rate in Labisia pumila Benth. under High CO2 and Nitrogen Fertilization, Molecules, vol.16, issue.1, pp.162-174, 2011.
DOI : 10.3390/molecules16010162

C. E. Jones and S. E. Hartley, A Protein Competition Model of Phenolic Allocation, Oikos, vol.86, issue.1, pp.27-44, 1999.
DOI : 10.2307/3546567

S. K. Lam, D. Chen, R. Norton, R. Armstrong, and A. R. Mosier, Nitrogen dynamics in grain crop and legume pasture systems under elevated atmospheric carbon dioxide concentration: A meta-analysis, Global Change Biology, vol.152, issue.9, pp.2853-2859, 2012.
DOI : 10.1007/s00442-007-0691-5

L. R. Larbar, . Bot, F. Bou-rgaud, C. Robin, and S. Adamowicz, Organ-specifie responses of tomate g rowth and phenolic metabolism to nitrate limitation, Plant Biol, 2012.

R. Larbar, K. M. Olsen, R. Slimestad, T. U>vdal, C. Bénard et al., Influence of repeated short-term nitrogen limitations on leaf phenolics metabolism in tomato, Phytochemistry, vol.77, pp.119-128, 2012.
DOI : 10.1016/j.phytochem.2012.02.004

W. Loomis, Growth-differentiation balance vs. carbohydrate-nitrogen ratio, Proc. Am Soc. Hort Sei, vol.29, pp.240-245, 1932.

T. Lllvdal, . K. Olsen, R. Slimestad, M. Verheul, and C. Üllo, Synergetic effects of nitrogen depletion, temperature, and light on the content of phenolic compounds and gene expression in leaves of tomato, Phytochemistry, vol.71, issue.5-6, pp.605-613, 2010.
DOI : 10.1016/j.phytochem.2009.12.014

A. Matros, S. Amme, B. Kettig, G. H. Buck-sorlin, U. Sonnewald et al., Growth at elevated CO2 concentrations leads to modified profiles of secondary metabolites in tobacco cv. SamsunNN and to increased resistance against infection with potato virus Y, Plant, Cell and Environment, vol.134, issue.1, pp.126-137, 2006.
DOI : 10.1146/annurev.phyto.27.1.143

D. Mckey, Adaptive Patterns in Alkaloid Physiology, The American Naturalist, vol.108, issue.961, pp.305-320, 1974.
DOI : 10.1086/282909

. D. Mckey, Distribution ofsecondary compounds within plants Herbivores: Their Interactio n w it h Secondary Plant Metabolites, pp.55-134, 1979.

V. Minden and M. Kleyer, Testing the effect-response framework: key response and effect traits determining above-ground biomass of salt marshes, Journal of Vegetation Science, vol.428, issue.2, pp.387-401, 2011.
DOI : 10.1038/nature02403

K. Mittelstrass, D. Treutter, M. Plessl, W. Helier, and E. F. Elstner, Modification of primary and secondary metabolism of potato plants by nitrogen application differentially a ffects resistance to Phyrophthora infestans and A/remaria solanL Plant Biol, pp.653-661, 2006.

L. Morrison and D. Stewart, Determination of lignin in t he presence of esterbound substituted cinnamic acids by a modified acetyl bromide procedure, J, 1995.

Y. Peng, K. Niklas, and S. Sun, The relationship between relative growth rate and whole-plant C : N : P stoichiometry in plant seedlings grown under nutrient-enriched conditions, Journal of Plant Ecology, vol.49, issue.8, pp.147-156, 2011.
DOI : 10.4319/lo.2004.49.4_part_2.1417

D. F. Rhoades, Evolution of plant chemical defense against herbivores Herbivores: Their lnreracticn with Secondary Plant Metabolites, pp.4-54, 1979.

C. Ruelas, M. E. Tiznado-hernândez, S. Estrada, A. O. Robles-burguer, M. R. Troncoso-rojas et al., Changes in phenolic acid content d uring a!oemaria altemata infection in tomate fruit j, pp.36-244, 2006.

J. Sardans, The C:N:P stoichiometry of organisms and ecosystems in a changing world: A review and perspectives, Perspectives in Plant Ecology, Evolution and Systematics, vol.14, issue.1, pp.33-47
DOI : 10.1016/j.ppees.2011.08.002

M. Schadler, M. Roeder, R. Bra-ndi, and D. Matthies, , nutrient availability and plant species on a generalist invertebrate herbivore, Global Change Biology, vol.26, issue.5, pp.1005-101, 2007.
DOI : 10.1046/j.1365-2486.2002.00497.x

W. R. Scheible, A. Gonzalez-fontes, M. Lauerer, B. Mulle-r-rober, M. Caboche et al., Nitrate Acts as a Signal to Induce Organic Acid Metabolism and Repress Starch Metabolism in Tobacco, THE PLANT CELL ONLINE, vol.9, issue.5, pp.783-798, 1997.
DOI : 10.1105/tpc.9.5.783

N. Stamp, can the growt h-different iation balance hypothesis be tested rigorously?, Forum, vol.107, pp.439-448, 2004.

. M. Stitt and A. Krapp, The interaction between elevated carbon dioxide and nitrogen nutrition: the physiological and molecular background, Plant, Cell and Environment, vol.23, issue.6, pp.583-621, 1999.
DOI : 10.1071/PP9960045

URL : https://hal.archives-ouvertes.fr/hal-01608545

J. Tuoml, Toward integration of plant defense theories, Trends EcoL Evol, vol.7, pp.365-367, 1992.

H. D. Van-etren, J. W. Mansfield, J. A. Bailey, and E. E. Far-mer, Two classes of plant antibiotics: Phytoalexins versus "Phytoanticipins, pp.91-102, 1994.

C. P. Vance, T. K. Kirk, and R. T. Sherwood, Lignification as a Mechanism of Disease Resistance, Annual Review of Phytopathology, vol.18, issue.1, pp.259-288, 1980.
DOI : 10.1146/annurev.py.18.090180.001355

A. Zangerl and M. R. Berenbaum, Plant Chemistry, Insect Adaptations to Plant Chemistry, and Host Plant Utilization Patterns, Ecology, vol.74, issue.1, pp.47-54, 1993.
DOI : 10.2307/1939500

A. Zobel and S. A. Brown, Seasonal changes of furanocoumarin concentrations in leaves ofHeracleum lanatum, Journal of Chemical Ecology, vol.16, issue.5, pp.1623-1634, 1990.
DOI : 10.1016/0165-1110(76)90012-9

P. Zornoza and . Sanz, Analysis of the interaction between light and the type of nitrogen nutrition in tomate plants, pp.389-398, 1991.

. Kaempferol-rutinoside-tomatine, 22) petite galerie 0.038 (±0.0078) 0.016 (±0.0053) 16.3 (±0.53) 10.6 (±1.79) grande galerie 0