, Rubus fruticosus: responses of growth, water status and ?13C composition, New Phytologist, vol.151, pp.427-435

M. N. Fotelli, H. Rennenberg, and A. Gessler, Effects of Drought on the Competitive Interference of an Early Successional Species (Rubus fruticosus) on Fagus sylvatica L. Seedlings: 15N Uptake and Partitioning, vol.4, pp.311-320, 2002.

M. N. Fotelli, M. Rienks, H. Rennenberg, and A. Gessler, Climate and forest management affect 15N-uptake, N balance and biomass of European beech seedlings, Trees -Structure and Function, vol.18, pp.157-166, 2004.
DOI : 10.1007/s00468-003-0289-4

M. N. Fotelli, M. Nahm, and K. Radoglou, Seasonal and interannual ecophysiological responses of beech (Fagus sylvatica) at its south-eastern distribution limit in Europe, Forest Ecology and Management, vol.257, pp.1157-1164, 2009.
DOI : 10.1016/j.foreco.2008.11.026

J. F. Franklin, H. H. Shugart, and M. E. Harmon, Tree Death as an Ecological Process, BioScience, vol.37, pp.550-556, 1987.
DOI : 10.2307/1310665

L. E. Frelich, The disturbance regime and its components. Forest Dynamics and Disturbance Regimes, pp.15-43, 2002.
DOI : 10.1017/cbo9780511542046.003

A. Gallé and U. Feller, Changes of photosynthetic traits in beech under severe drought stress and during recovery, Physiologia Plantarum, vol.131, pp.412-421, 2007.

D. A. Galvez, S. M. Landhäusser, and M. T. Tyree, Root carbon reserve dynamics in aspen seedlings: does simulated drought reserve limitation?, Tree Physiology, vol.31, pp.250-257, 2011.
DOI : 10.1093/treephys/tpr012

URL : https://academic.oup.com/treephys/article-pdf/31/3/250/4605393/tpr012.pdf

N. Garcia-forner, H. D. Adams, and S. Sevanto, Responses of two semiarid conifer tree species to reduced precipitation and warming reveal new perspectives for stomatal regulation, Plant, Cell & Environment, vol.152, pp.412-418, 2015.
DOI : 10.1111/pce.12588

URL : https://ddd.uab.cat/pub/artpub/2016/141507/placelenv_a2016v39n1.pdf

P. Gasson, Some implications of anatomical variations in the wood of pedunculate oak (Quercuc robur L.), including comparison with common beech, Fagus sylvatica L.). IAWA Bull, vol.8, pp.149-166, 1987.

J. B. Gaudinski, M. S. Torn, W. J. Riley, C. Swanston, S. E. Trumbore et al., Use of stored carbon reserves in growth of temperate tree roots and leaf buds: analyses using radiocarbon measurements and modeling, Global Change Biology, vol.15, pp.992-1014, 2009.

M. L. Gaylord, T. E. Kolb, W. T. Pockman, J. A. Plaut, E. A. Yepez et al., Drought predisposes piñon-juniper woodlands to insect attacks and mortality, New Phytologist, vol.198, pp.567-578, 2013.
DOI : 10.1111/nph.12174

URL : https://nph.onlinelibrary.wiley.com/doi/pdf/10.1111/nph.12174

M. L. Gaylord, T. E. Kolb, and N. G. Mcdowell, Mechanisms of pinon pine mortality after severe drought: a retrospective study of mature trees, Tree Physiology, vol.35, pp.806-816, 2015.

H. Genet, N. Bréda, and E. Dufrêne, Age-related variation in carbon allocation at tree and stand scales in beech (Fagus sylvatica L.) and sessile oak (Quercus petraea (Matt.) Liebl.) using a chronosequence approach, Tree Physiology, vol.30, pp.177-192, 2009.
DOI : 10.1093/treephys/tpp105

URL : https://academic.oup.com/treephys/article-pdf/30/2/177/4787482/tpp105.pdf

B. Gérard and N. Bréda, Radial distribution of carbohydrate reserves in the trunk of declining European beech trees (Fagus sylvatica L.), Annals of Forest Science, vol.71, pp.675-682, 2012.

G. Grassi, P. Millard, R. Wendler, G. Minotta, and M. Tagliavini, Measurement of xylem sap amino acid concentrations in conjunction with whole tree transpiration estimates spring N re-mobilization by cherry, Prunus avium L.) trees. Plant, Cell and Environment, vol.25, pp.1689-1699, 2002.

G. Grassi, P. Millard, P. Gioacchini, and M. Tagliavini, Recycling of nitrogen in the xylem of Prunus avium trees starts when spring remobilization of internal reserves declines, Tree physiology, vol.23, pp.1061-1068, 2003.

J. J. Green, L. A. Dawson, J. Proctor, E. I. Duff, and D. A. Elston, Fine root dynamics in a tropical rain forest is influenced by rainfall, Plant Soil, vol.276, pp.23-32, 2005.

S. Greenwood, P. Ruiz-benito, and J. Martínez-vilalta, Tree mortality across biomes is promoted by drought intensity, lower wood density and higher specific leaf area, Ecology Letters, vol.20, pp.539-553, 2017.

P. L. Gregersen, P. B. Holm, and K. Krupinska, Leaf senescence and nutrient remobilisation in barley and wheat, Plant Biology, vol.10, pp.37-49, 2008.

A. Gruber, D. Pirkebner, C. Florian, and W. Oberhuber, No evidence for depletion of carbohydrate pools in Scots pine (Pinus sylvestris L.) under drought stress, Plant Biology, vol.14, pp.142-148, 2012.

B. D. Gruber, R. F. Giehl, S. Friedel, and N. Von-wirén, Plasticity of the Arabidopsis root system under nutrient deficiencies, Plant physiology, vol.163, pp.161-79, 2013.

L. Gu, S. G. Pallardy, K. P. Hosman, and Y. Sun, Predictors and mechanisms of the drought-influenced mortality of tree species along the isohydric to anisohydic continuum in a decade-long study of a central US temperate forest, Biogeoscience Discussion, vol.12, pp.1285-1325, 2015.

S. Guak, D. Neilsen, P. Millard, R. Wendler, and G. H. Neilsen, Determining the role of N remobilization for growth of apple (Malus domestica Borkh.) trees by measuring xylem-sap N flux, Journal of Experimental Botany, vol.54, pp.2121-2131, 2003.

E. J. Gustafson, Applicability of predictive models of drought-induced tree mortality between the midwest and northeast United States, Forest Science, vol.60, pp.327-334, 2014.

U. Hacke and J. J. Sauter, Xylem dysfunction during winter and recovery of hydraulic conductivity in diffuse-porous and ring-porous trees, Oecologia, vol.105, pp.425-439, 1996.

Z. Hai-yang, L. Xiao-tao, and H. Hartmann, Foliar nutrient resorption differs between arbuscular mycorrhizal and ectomycorrhizal trees at local and global scales, Global Ecology and Biogeography, vol.27, 2018.

J. Hansen and I. Møller, Percolation of starch and soluble carbohydrates from plant tissue for quantitative determination with anthrone, Analytical Biochemistry, vol.68, pp.87-94, 1975.

P. J. Hanson and J. F. Weltzin, Drought disturbance from climate change: response of United States forests, The Science of the Total Environnement, vol.262, pp.205-220, 2000.

D. A. Herms and W. J. Mattson, The dilemma of plants: to grow or defend, The Quarterly Review of Biology, vol.67, pp.282-335, 1992.

K. A. Harrison, R. Bol, and R. D. Bardgett, Do plant species with different growth strategies vary in their ability to compete with soil microbes for chemical forms of nitrogen?, Soil Biology and Biochemistry, vol.40, pp.228-237, 2008.

T. Hölttä, H. Cochard, E. Nikinmaa, and M. Mencuccini, Capacitive effect of cavitation in xylem conduits: Results from a dynamic model, Plant, Cell and Environment, vol.32, pp.10-21, 2009.

T. C. Hsiao, E. Acevedo, E. Fereres, and D. W. Henderson, Water stress, growth and osmotic adjustment, Philosophical Transactions of the Royal Society London B, vol.273, pp.479-500, 1976.

T. C. Hsiao, Plant responses to water stress, Annual Review of Plant Biology, vol.24, pp.519-570, 1973.

C. Huntingford, P. Zelazowski, D. Galbraith, L. M. Mercado, S. Sitch et al., Simulated resilience of tropical rainforests to CO2-induced climate change, Nature Geoscience, vol.6, pp.268-273, 2013.

R. Hyvonen, G. I. Agren, and S. Linder, The likely impact of elevated [CO2], nitrogen deposition, increased temperature and management on carbon sequestration in temperate and boreal forest ecosystems: a literature review, New Phytologist, vol.173, pp.463-480, 2007.

, Inventaire Forestier National (IFN), 2013.

D. Stocker, T. F. Qin, G. Plattner, M. Tignor, S. K. Allen et al., Contribution of Working Group I to the fifth assessment report of the intergovernmental panel on climate change, pp.159-254

. Ipcc, Global Warming of 1.5°C. Summary for Policymakers, 2018.

R. B. Jackson, H. A. Mooney, and E. D. Schulze, A global budget for fine root biomass, surface area, and nutrient contents, Plant National Academic Science USA, vol.94, pp.7362-7366, 1997.

J. Jacquet, Impacts des défoliations de la processionnaire du pin (Thaumetopoea pityocampa) sur la croissance du pin maritime (Pinus pinaster), École Doctorale: Sciences et Environments Ph, p.151, 2012.

A. Janská, P. Mar?ík, S. Zelenková, and J. Ovesná, Cold stress and acclimation -what is important for metabolic adjustment, Plant Biology, vol.12, pp.395-405, 2010.

S. Jarvis and J. Macduff, Nitrate nutrition of grasses from steady-state supplies in flowing solution culture following nitrate deprivation and/or defoliation. I recovery of uptake and growth and their interactions, Journal of Experimental Botany, vol.40, pp.965-975, 1989.

S. N. Johnson, M. Erb, and S. E. Hartley, Roots under attack: Contrasting plant responses to below-and aboveground insect herbivory, New Phytologist, vol.210, pp.413-418, 2016.

M. O. Jordan, R. Wendler, and P. Millard, Autumnal N storage determines the spring growth, N uptake and N internal cycling of young peach trees, Trees-Structure and Function, vol.26, pp.393-404, 2012.

M. O. Jordan, 2015. C depletion and tree dieback in young peach trees : a possible consequence of N shortage ?, Annals of forest science, vol.5, p.537
URL : https://hal.archives-ouvertes.fr/hal-01595774

J. Jourdan, Variations saisonnières de la morphogenèse et de la croissance des systèmes aériens et souterrains chez le peuplier, Leurs relations avec les glucides et les transferts minéraux, p.160, 1980.

A. S. Jump, C. Mátyás, and J. Peñuelas, The altitude-for-latitude disparity in the range retractions of woody species, Trends in Ecology and Evolution, vol.24, pp.694-701, 2009.

S. M. Kang, K. C. Ko, and J. S. Titus, Mobilization and metabolism of pro-tein and soluble nitrogen during spring growth of apple-trees, Journal of the American Society for Horticultural Science, vol.107, pp.209-213, 1982.

H. Kawaletz, I. Mölder, P. Annighöfer, A. Terwei, S. Zerbe et al., Pot experiments with woody species a review, Forestry, vol.87, pp.482-491, 2014.

S. G. Keel and C. Schädel, Expanding leaves of mature deciduous forest trees rapidly become autotrophic, Tree Physiology, vol.30, pp.1253-1259, 2010.

T. Kiba, T. Kudo, M. Kojima, and H. Sakakibara, Hormonal control of nitrogen acquisition: roles of auxin, abscisic acid, and cytokinin, Journal of Experimental Botany, vol.62, pp.1399-1409, 2011.

T. Kiba and A. Krapp, Plant nitrogen acquisition under low availability: regulation of uptake and root architecture, Plant and Cell Physiology, vol.57, pp.707-714, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01531685

K. T. Killingbeck, Nutrients in senesced leaves: Keys to the search for potential resorption and resorption proficiency, Ecology, vol.77, pp.1716-1727, 1996.

A. W. King, W. M. Post, and S. D. Wullschleger, The potential response of terrestrial car-bon storage to changes in climate and atmospheric CO2, Climatic Change, vol.35, pp.199-227, 1997.

I. Klein and S. A. Weinbaum, Foliar application ofurea to olive: translocation of urea nitrogen as influenced by sink demand and nitrogen deficiency, Journal of the American Society for Horticultural Science, vol.109, pp.356-360, 1984.

T. Klein, G. Hoch, D. Yakir, and C. Körner, Drought stress, growth and nonstructural carbohydrate dynamics of pine trees in a semi-arid forest, Tree Physiology, vol.34, pp.981-992, 2014.

M. Knoblauch and W. S. Peters, Münch, morphology, microfluidics -our structural problem with the phloem: Münch, morphology, microfluidics, Plant Cell Environment, vol.33, pp.1439-1452, 2010.

M. Knoche, P. D. Petracek, M. J. Bukovac, and W. E. Shafer, Urea Penetration of Isolated Tomato Fruit Cuticles, Journal of the American Society for Horticultural Science, vol.119, pp.761-764, 1994.

T. E. Kolb, K. A. Dodds, and K. M. Clancy, Effect of western spruce budworm defoliation on the physiology and growth of potted Douglas-fir seedlings, Forest Science, vol.45, pp.281-291, 1999.

C. Körner, Carbon limitation in trees, Journal of Ecology, vol.91, pp.4-17, 2003.

K. R. Kosola, D. I. Dickmann, E. A. Paul, and D. Parry, Repeated insect defoliation effects on growth, nitrogen acquisition, carbohydrates, and root demography of poplars, Oecologia, vol.129, pp.65-74, 2001.

T. T. Kozlowski and S. G. Pallardy, Physiology of woody plants, 1997.

P. J. Kramer and T. T. Kozlowski, Physiology of woody plants, 1979.

K. Kramer, B. Degen, J. Buschbom, T. Hickler, W. Thuller et al.,

, Modelling exploration of the future of European beech (Fagus sylvatica L.) under climate change -range, abundance, genetic diversity and adaptive response, Forest Ecology and Management, vol.259, pp.2213-2222

G. H. Krause, U. Arndf, and C. J. Brandt, Forest decline in Europe: development and possible causes, Water, Air and Soil Pollution, vol.31, pp.647-668, 1986.

S. Krause, K. Raffa, and M. Wagner, Tree responses to stress: a role in sawfly outbreaks, Sawfly life history adaptations to woody plants, pp.211-227, 1993.

J. Kreuzwieser and A. Gessler, Global climate change and tree nutrition: Influence of water availability, Tree Physiology, vol.30, pp.1221-1234, 2010.

S. Kulac, P. Nzokou, D. Guney, B. M. Cregg, and I. Turna, Growth and physiological response of Fraser fir [Abies fraseri (Pursh) Poir.] seedlings to water stress: seasonal and diurnal variations in photosynthetic pigments and carbohydrate concentration, HortScience, vol.47, pp.1512-1519, 2012.

H. M. Kulman, Effects of insect defoliation on growth and mortality of trees, Annual Review of Entomology, vol.16, pp.289-324, 1971.

A. Lacointe, A. Kajji, F. A. Daudet, P. Archer, and J. S. Frossard, Mobilization of carbon reserves in young walnut trees, Acta Botanica Gallica, vol.140, pp.435-441, 1993.

H. M. Lam, K. T. Coschigano, I. C. Oliveira, R. Melo-oliveira, and G. M. Coruzzi, The molecular-genetics of nitrogen assimilation into amino acids in higher plants, Annual Review of Plant Physiology and Plant Molecular Biology, vol.47, pp.569-593, 1996.

M. Larter, S. Pfautsch, J. Domec, S. Trueba, N. Nagalingum et al., Aridity drove the evolution of extreme embolism resistance and the radiation of conifer genus Callitris, New Phytologist, vol.215, pp.97-112, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01606790

S. Lavorel, J. Lebreton, L. Maho, and Y. , Les mécanismes d'adaptation de la biodiversité aux changements climatiques et leurs limites, 2017.

F. Lebourgeois, N. Bréda, E. Ulrich, and A. Granier, Climate-tree-growth relationships of European beech (Fagus sylvatica L.) in the French Permanent Plot Network (RENECOFOR), Trees -Structure and Function, vol.19, pp.385-401, 2005.
URL : https://hal.archives-ouvertes.fr/hal-01831367

F. Lebourgeois, J. Pierrat, and V. Perez, Simulating phenological shifts in French temperate forests under two climatic change scenar-ios and four driving global circulationmodels, International Journal of Biometeorology, vol.54, pp.563-581, 2010.

D. Lemoine, H. Cochard, and A. Granier, Within crown variation in hydraulic architecture in beech (Fagus sylvatica L): evidence for a stomatal control of xylem embolism, Annals of Forest Science, vol.59, pp.19-27, 2002.
URL : https://hal.archives-ouvertes.fr/halsde-00153133

C. Leuschner, K. Backes, and D. Hertel, Drought responses at leaf, stem and fine root levels of competitive Fagus sylvatica L. and Quercus petraea (Matt.) Liebl. trees in dry and wet years, Forest Ecology and Management, vol.149, pp.33-46, 2001.
DOI : 10.1016/s0378-1127(00)00543-0

M. H. Li, G. Hoch, and C. Korner, Source/sink removal affects mobile carbohydrates in Pinus cembra at the Swiss treeline, Trees Structure and Function, vol.16, pp.331-337, 2002.

J. Li, T. L. Powell, T. J. Seiler, D. P. Johnson, and H. P. Anderson, Hurricane Frances on Florida scrub-oak ecosystem processes: defoliation, net CO2 exchange and interactions with elevated CO2, Global Change Biology, vol.13, pp.1101-1113, 2007.

P. O. Lim, H. J. Kim, and H. G. Nam, Leaf senescence, Annual Review of Plant Biology, vol.58, pp.115-136, 2007.

M. J. Linton, J. S. Sperry, and D. G. Williams, Limits to water transport in Juniperus osteosperma and Pinus edulis: implications for drought tolerance and regulation of transpiration, Functional Ecology, vol.12, pp.906-911, 1998.

Y. Liu, X. Chen, and A. Ouyang, Nondestructive determination of pear internal quality indices by visible and near-infrared spectrometry, LWT -Food Science and Technology, vol.41, pp.1720-1725, 2008.
DOI : 10.1016/j.lwt.2007.10.017

B. Liu, J. X. He, F. J. Zeng, J. Q. Lei, and S. K. Arndt, Life span and structure of ephemeral root modules of different functional groups from a desert system, New Phytologist, vol.211, pp.103-112, 2016.

J. Liu, M. Arend, and W. Yang, Effects of drought on leaf carbon source and growth of European beech are modulated by soil type, vol.7, pp.1-9, 2017.

G. Llácer, Hosts and symptoms of plum pox virus: Herbaceous hosts, EPPO Bulletin, vol.36, pp.227-228, 2006.

F. Lloret, J. Peñuelas, and M. Estiarte, Experimental evidence of reduced diversity of seedlings due to climate modification in a Mediterranean-type community, Global Change Biology, vol.10, pp.248-258, 2004.

F. Lloret, A. Escudero, J. M. Iriondo, J. Mart?nez-vilalta, and F. Valladares, Extreme climatic events and vegetation: the role of stabilizing processes, Global Change Biology, vol.18, pp.797-805, 2012.

J. Lopez-bucio, A. Cruz-ramirez, and L. Herrera-estrella, The role of nutrient availability in regulating root architecture, Current Opinion in Plant Biology, vol.6, pp.280-287, 2003.

M. Lorenz and G. Becher, Forest Condition in Europe, Technical Report of ICP Forests. Work Report of the Thunen Institute for World Forestry, 2012.

D. Loustau, Forests, carbon cycle and climate change, QUAE, vol.350, p.pp, 2010.

D. Loustau and A. Bosc, Contraintes climatiques sur les forêts, Innovations Agronomiques, vol.18, pp.71-86, 2012.

G. M. Lovett, L. M. Christenson, P. M. Groffman, C. G. Jones, J. E. Hart et al., Insect defoliation and nitrogen cycling in forests, BioScience, vol.52, pp.335-341, 2002.

X. Lu, G. T. Freschet, D. F. Flynn, and X. Han, Plasticity in leaf and stem nutrient resorption proficiency potentially reinforces plant-soil feedbacks and microscale heterogeneity in a semi-arid grassland, Journal of Ecology, vol.100, pp.144-150, 2012.

J. Luo, H. Li, T. Liu, A. Polle, C. Peng et al., Nitrogen metabolism of two contrasting poplar species during acclimation to limiting nitrogen availability, Journal of Experimental Botany, vol.64, pp.4207-4224, 2013.

J. M. Lynch and J. M. Whipps, Substrate flow in the rhizosphere, Plant Soil, vol.129, pp.1-10, 1990.

J. H. Macaduff, S. C. Jarvis, and A. Mosquera, Nitrate nutrition of grasses from steady-state supplies in flowing solution culture following nitrate deprivation and/or defoliation: II. Assimilation of NO3 and/or short-term effects on NO3 uptake, Journal of Experimental Botany, vol.40, pp.977-984, 1989.

F. Magnani and M. Borghetti, Interpretation of Seasonal-Changes of Xylem Embolism and Plant Hydraulic Resistance in Fagus-Sylvatica, Plant Cell and Environment, vol.18, pp.689-696, 1995.

A. J. Maguire and R. K. Kobe, Drought and shade deplete nonstructural carbohydrate reserves in seedlings of five temperate tree species, Ecology and Evolution, vol.5, pp.5711-5721, 2015.

D. Malaguti, P. Millard, R. Wendler, A. Hepburn, and M. Tagliavini, Translocation of amino acids in the xylem of apple (Malus domestica Borkh.) trees in spring as a consequence of both N remobilization and root uptake, Journal of experimental botany, vol.52, pp.1665-1671, 2001.

P. D. Manion, Tree disease concepts, 1981.

À. Manrique-alba, S. Sevanto, and H. D. Adams, Stem radial growth and water storage responses to heat and drought vary between conifers with differing hydraulic strategies, Plant Cell and Environment, vol.41, pp.1926-1934, 2018.

J. Martí-nez-vilalta, J. Piñol, and K. Beven, A hydraulic model to predict drought-induced mortality in woody plants: an application to climate change in the Mediterranean, Ecological Modelling, vol.155, pp.127-147, 2002.

J. Martínez-vilalta, F. Lloret, and D. D. Breshears, Drought-induced forest decline: causes, scope and implications, Biology Letters, vol.8, pp.689-691, 2012.

N. Martin-stpaul, S. Delzon, and H. Cochard, Plant resistance to drought depends on timely stomatal closure, Ecology Letters, vol.20, pp.1437-1447, 2017.

C. Masclaux-daubresse, F. Daniel-vedele, J. Dechorgnat, F. Chardon, L. Gaufichon et al., Nitrogen uptake, assimilation and remobilization in plants: Challenges for sustainable and productive agriculture, Annals of Botany, vol.105, pp.1141-1157, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01203920

M. L. Mccormack and D. L. Guo, Impacts of environmental factors on fine root lifespan, Frontiers in Plant Science, vol.5, p.205, 2014.

M. L. Mccormack, I. A. Dickie, and D. M. Eissenstat, Redefining fine roots improves understanding of below-ground contributions to terrestrial biosphere processes, New Phytologist, vol.207, pp.505-518, 2015.

N. Mcdowell, W. T. Pockman, and C. D. Allen, Mechanisms of plant survival and mortality during drought: Why do some plants survive while others succumb to drought, New Phytologist, vol.178, pp.719-739, 2008.

N. G. Mcdowell, M. G. Ryan, M. J. Zeppel, and D. T. Tissue, Improving our knowledge of drought-induced forest mortality through experiments, observations, and modeling, New Phytologist, vol.200, pp.289-293, 2013.

S. B. Mclaughlin, R. K. Mcconathy, R. L. Barnes, and N. T. Edwards, Seasonal changes in energy allocation by white oak (Quercus alba), Canadian Journal of Forest Research, vol.10, pp.379-388, 1980.

L. Mei, Y. Xiong, J. Gu, and Z. Wang, Whole -tree dynamics of non-structural carbohydrate and nitrogen pools across different seasons and in response to girdling in two temperate trees, vol.177, pp.333-344, 2015.

F. C. Meinzer, D. M. Johnson, B. Lachenbruch, K. A. Mcculloh, and D. R. Woodruff, , 2009.

, Xylem hydraulic safety margins in woody plants: coordination of stomatal control of xylem tension with hydraulic capacitance, Functional Ecology, vol.23, pp.922-930

F. C. Meinzer, Stomatal kinetics and photosynthetic gas exchange along a continuum of iso-to anisohydric regulation of plant water status, Plant Cell and Environment, vol.40, pp.1618-1628, 2016.

P. Meir, M. Mencuccini, and R. C. Dewar, Drought-related tree mortality: addressing the gaps in understanding and prediction, New Phytologist, vol.207, pp.28-33, 2015.

M. Mencuccini, Temporal scales for the coordination of tree carbon and water economies during droughts, Tree Physiology, vol.34, pp.439-442, 2014.

A. Michelot, S. Simard, C. Rathgeber, E. Dufrêne, and C. Damesin, Comparing the intra-annual wood formation of three European species (Fagus sylvatica, Quercus petraea and Pinus sylvestris) as related to leaf phenology and non-structural carbohydrate dynamics, Tree Physiology, vol.32, pp.1033-1045, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01004139

M. Migliavacca, O. Sonnentag, and T. F. Keenan, On the uncertainty of phenological responses to climate change, and implications for a terrestrial biosphere model, Biogeosciences, vol.9, pp.2063-2083, 2012.

P. Millard and G. H. Neilsen, The influence of nitrogen supply on the uptake atid remohilisation of stored N for the seasonal growth of apple ttees, Annals of Botany, vol.63, p.301, 1989.

P. Millard and M. F. Proe, Leaf demography and the seasonal internal cycling of nitrogen in sycamore (Acer pseudoplatanus L.) seedlings in relation to nitrogen supply, New Phytologist, vol.117, pp.587-596, 1991.

P. Millard, A. Hester, R. Wendler, and G. Baillie, Interspecific defoliation responses of trees depend on sites of winter nitrogen storage, Functional Ecology, vol.15, pp.535-543, 2001.

P. Millard, M. Sommerkorn, and G. A. Grelet, Environmental change and carbon limitation in trees: a biochemical, ecophysiological and ecosystem appraisal, New Phytologist, vol.175, pp.11-28, 2007.

P. Millard and G. A. Grelet, Nitrogen storage and remobilization by trees: Ecophysiological relevance in a changing world, Tree Physiology, vol.30, pp.1083-1095, 2010.

P. Millard, The accumulation and storage of nitrogen by herbaceous plants, Plant Cell and Environment, vol.11, pp.1-8, 1988.

P. Millard, Ecophysiology of the internal cycling of nitrogen for tree growth, Zeitschrift für Pflanzenernährung und Bodenkunde, vol.159, pp.1-10, 1996.

J. Millett, P. Millard, A. J. Hester, and A. J. Mcdonald, Do competition and herbivory alter the internal nitrogen dynamics of birch saplings?, New Phytologist, vol.168, pp.413-422, 2005.

P. J. Mitchell, A. P. O'grady, D. T. Tissue, D. A. White, M. L. Ottenschlaeger et al., Drought response strategies define the relative con-tributions of hydraulic dysfunction and carbohydrate depletion during tree mortality, New Phytologist, vol.197, pp.862-872, 2013.

H. A. Mooney and R. I. Hays, Carbohydrate storage cycles in two Californian Mediterraneanclimate trees, Flora, vol.162, pp.295-304, 1973.

E. Moran, J. Lauder, C. Musser, A. Stathos, and M. Shu, The genetics of drought tolerance in conifers, New Phytologist, vol.216, pp.1034-1048, 2017.

J. Muhr, A. Angert, R. I. Negrón-juárez, W. Muñoz, G. Kraemer et al., Carbon dioxide emitted from live stems of tropical trees is several years old, Tree Physiology, vol.33, pp.743-752, 2013.

B. Muller, F. Pantin, M. Génard, O. Turc, S. Freixes et al., Water deficits uncouple growth from photosynthesis, increase C content, and modify the relationships between C and growth in sink organs, Journal of Experimental Botany, vol.62, pp.1715-1729, 2011.

E. Münch, Die Stoffbewegungen in der Pflanze, 1930.

E. Nabeshima, M. Murakami, and T. Hiura, Effects of herbivory and light conditions on induced defense in Quercus crispula, Journal of Plant Research, vol.114, pp.403-409, 2001.

G. J. Nabuurs, M. J. Schelhaas, G. M. Mohren, and C. B. Field, Temporal evolution of the European forest sector carbon sink from 1950 to 1999, Global Change Biology, vol.9, pp.152-160, 2003.

L. M. Nageleisen, D. Piou, F. X. Saintonge, and P. Riou-nivert, La santé des forets, Maladies, insectes, accidents climatiques ? diagnostic et prévention, 2010.

M. Nahm, A. Matzarakis, H. Rennenberg, and A. Gessler, Seasonal courses of key parameters of nitrogen, carbon and water balance in European beech (Fagus sylvatica L.) grown on four different study sites along a European North-South climate gradient during the 2003 drought, Trees -Structure and Function, vol.21, pp.79-92, 2007.

R. K. Nair, M. P. Perks, and A. Weatherall, Does canopy nitrogen uptake enhance carbon sequestration by trees?, Global Change Biology, vol.22, pp.875-888, 2015.

A. Nardini, M. Battistuzzo, and T. Savi, Shoot desiccation and hydraulic failure in temperate woody angiosperms during an extreme summer drought, The New Phytologist, vol.200, pp.322-329, 2013.

D. Neilsen, P. Millard, G. H. Neilsen, and E. J. Hogue, Sources of N for leaf growth in a high-density apple (Malus domestica) orchard irrigated with ammonium nitrate solution, Tree physiology, vol.17, pp.733-739, 1997.

E. A. Nelson and R. E. Dickson, Accumulation of food reserves in cottonwood stems during dormancy induction, Canadian Journal of Forest Research, vol.11, pp.145-154, 1981.

S. Netherer, B. Matthews, and K. Katzensteiner, Do water-limiting conditions predispose Norway spruce to bark beetle attack, New Phytologist, vol.205, pp.1128-1141, 2015.

E. Nikinmaa, T. Hölttä, P. Hari, P. Kolari, A. Mäkelä et al., Assimilate transport in phloem sets conditions for leaf gas exchange, Plant, Cell and Environment, vol.36, pp.655-669, 2013.

M. Nolf, R. Lopez, and J. M. Peters, Visualization of xylem embolism by X-ray microtomography: a direct test against hydraulic measurements, New Phytologist, vol.214, pp.890-898, 2017.

M. J. O'brien, D. F. Burslem, A. Caduff, J. Tay, and A. Hector, Contrasting nonstructural carbohydrate dynamics of tropical tree seedlings under water deficit and variability, New Phytologist, vol.205, pp.1083-1094, 2015.

M. J. O'brien, B. M. Engelbrecht, J. Joswig, G. Pereyra, B. Schuldt et al., A synthesis of tree functional traits related to drought-induced mortality in forests across climatic zones, Journal of Applied Ecology, vol.54, pp.1669-1686, 2017.

B. T. O'kenney, M. J. Hennerty, and J. S. Titus, Changes in the nitrogen reserves of apple shoots during the dormant season, The Journal of Horticultural Science and Biotechnology, vol.50, pp.321-329, 1975.

E. Ögren, Maintenance respiration correlates with sugar but not nitrogen concentration in dormant plants, Physiology Plant, vol.108, pp.295-299, 2000.

J. Oliva, J. Stenlid, and J. Martínez-vilalta, The effect of fungal pathogens on the water and carbon economy of trees: implications for drought-induced mortality included in theoretical models for drought induced mortality, New Phytologist, vol.203, pp.1028-1035, 2014.

M. M. Ortega-loeza, R. Salgado-garciglia, C. Gomez-alonso, A. Diaz, and I. , Acclimatization of the endangered mexican epiphytic orchid, Laelia speciosa, European Journal of Environmental Sciences, vol.1, pp.48-54, 2011.

A. Ourry, J. H. Macduff, J. J. Volenec, and J. P. Gaudillere, Nitrogen traffic during plant growth and development, Morot-Gaudry JF, Lea P Plant nitrogen, pp.255-273, 2001.

R. Ozolincius and V. Stakenas, Tree crown defoliation: influencing factors, Baltic Forestry, vol.2, pp.48-55, 1996.

S. Palacio, A. J. Hester, M. Maestro, and P. Millard, Browsed Betula pubescens trees are not carbon-limited, Functional Ecology, vol.22, pp.808-815, 2008.

S. Palacio, R. Hernández, M. Maestro-martínez, and J. J. Camarero, Fast replenishment of initial carbon stores after defoliation by the pine processionary moth and its relationship to the re-growth ability of trees, Trees -Structure and Function, vol.26, pp.1627-1640, 2012.

S. Palacio, G. Hoch, A. Sala, C. Körner, and P. Millard, Does carbon storage limit tree growth, New Phytologist, vol.201, pp.1096-1100, 2014.
DOI : 10.1111/nph.12602

URL : https://digital.csic.es/bitstream/10261/91703/1/Palacio%20et%20al_C%20limitation%20in%20trees_submitted%20New%20Phytol_R1_Final_clean.pdf

S. Palacio, J. J. Camarero, M. Maestro, A. Q. Alla, E. Lahoz et al., Are storage and tree growth related? Seasonal nutrient and carbohydrate dynamics in evergreen and deciduous Mediterranean oaks, Trees -Structure and Function, vol.32, pp.1-14, 2018.
DOI : 10.1007/s00468-018-1671-6

R. E. Pangle, J. M. Limousin, and . Plaut, Prolonged experimental drought reduces plant hydraulic conductance and transpiration and increases mortality in a piñon-juniper woodland, Ecology and evolution, vol.5, pp.1618-1623, 2015.
DOI : 10.1002/ece3.1422

URL : https://onlinelibrary.wiley.com/doi/pdf/10.1002/ece3.1422

F. Pantin, A. Fanciullino, C. Massonnet, M. Dauzat, T. Simonneau et al., , 2013.

, Buffering growth variations against water deficits through timely carbon usage, Frontiers in Plant Science, vol.4, pp.1-11

A. Parsons, E. Leafe, B. Collett, and W. Stiles, The physiology of grass produc-tion under grazing. I. Characteristics of leaf and canopy photosynthesis of continuously-grazed swards, Journal of Applied Ecology, vol.20, pp.117-126, 1983.

A. Patakas, N. Nikolaou, E. Zioziou, K. Radoglou, and B. Noitsakis, The role of organic solute and ion accumulation in osmotic adjustment in drought-stressed grapevines, Plant Science, vol.163, pp.361-367, 2002.

M. S. Peek, Explaining variation in fine root life span, Progress in Botany, pp.382-398, 2007.
DOI : 10.1007/978-3-540-36832-8_17

F. W. Penning-de-vries, The Cost of Maintenance Processes in Plant Cells, Annals of Botany, vol.39, pp.77-92, 1975.

J. J. Peñuelas, T. Rosas, L. Galiano, R. Ogaya, J. J. Peñuelas et al., Dynamics of non-structural carbohydrates in three Mediterranean woody species following long-term experimental drought, Frontiers in Plant Science, vol.4, pp.1-16, 2013.

A. D. Peuke and H. Rennenberg, 2004 Carbon, nitrogen, phosphorus, and sulphur concentration and partitioning in beech ecotypes (Fagus sylvatica L.): phosphorus most affected by drought, Trees -Structure and Function, vol.18, pp.639-648
DOI : 10.1007/s00468-004-0335-x

D. Peuke, C. Schraml, W. Hartung, and H. Rennenberg, Identification of drought-sensitive beech ecotypes by physiological parameters, New Phytologist, vol.154, pp.373-387, 2002.
DOI : 10.1046/j.1469-8137.2002.00400.x

H. Piene and C. H. Little, Spruce budworm defoliation and growth loss in young balsam fir: artificial defoliation of potted trees, Canadian Journal of Forest Research, vol.20, pp.902-909, 1990.

E. A. Pinkard, M. Battaglia, S. Roxburgh, and A. P. O'grady, Estimating forest net primary production under changing climate: adding pests into the equation, Tree physiology, vol.31, pp.686-699, 2011.
DOI : 10.1093/treephys/tpr054

G. Piovesan, F. Biondi, A. Filippo, A. Alessandrini, and M. Maugeri, Drought-driven growth reduction in old beech (Fagus sylvatica L.) forests of the central Apennines, Italy. Global Change Biology, vol.14, pp.1-17, 2008.

F. I. Piper, Drought induces opposite changes in the concentra-tion of non-structural carbohydrates of two evergreen Nothofagus species of differential drought resistance, Annals of Forest Science, vol.68, pp.415-424, 2011.

F. I. Piper, Patterns of carbon storage in relation to shade tolerance in southern South American species, American Journal of Botany, vol.102, pp.1442-1452, 2015.

F. I. Piper and A. Fajardo, Carbon dynamics of Acer pseudoplatanus seedlings under drought and complete darkness, Tree Physiology, vol.36, pp.1400-1408, 2016.
DOI : 10.1093/treephys/tpw063

F. I. Piper and A. Fajardo, Foliar habit, tolerance to defoliation and their link to carbon and nitrogen storage, Journal of Ecology, vol.102, pp.1101-1111, 2014.
DOI : 10.1111/1365-2745.12284

URL : https://besjournals.onlinelibrary.wiley.com/doi/pdf/10.1111/1365-2745.12284

J. Pittermann, The evolution of water transport in plants: an integrated approach, Geobiology, vol.8, pp.112-139, 2010.

H. Poorter, F. Fiorani, M. Stitt, U. Schurr, A. Finck et al., The art of growing plants for experimental purposes: a practical guide for the plant biologist, Functional Plant Biology, vol.39, pp.821-838, 2012.

R. B. Pratt, A. L. Jacobsen, R. Mohla, F. W. Ewers, and S. D. Davis, Linkage between water stress tolerance and life history type in seedlings of nine chaparral species (Rhamnaceae), Journal of Ecology, vol.96, pp.1252-1265, 2008.

C. Preece, G. Farré-armengol, J. Llusià, and J. Peñuelas, Thirsty tree roots exude more carbon, Tree Physiology, vol.38, pp.690-695, 2018.
DOI : 10.1093/treephys/tpx163

E. Puri, G. Hoch, and C. Korner, Defoliation reduces growth but not carbon reserves in Mediterranean Pinus pinaster trees, Trees, vol.29, pp.1187-1196, 2015.
DOI : 10.1007/s00468-015-1199-y

A. G. Quentin, A. P. O'grady, C. L. Beadle, D. Worledge, and E. A. Pinkard, Responses of transpiration and canopy conductance to partial defoliation of Eucalyptus globulus trees, Agricultural and Forest Meteorology, vol.151, pp.356-364, 2011.

A. G. Quentin, E. A. Pinkard, and M. G. Ryan, Non-structural carbohydrates in woody plants compared among laboratories, Tree Physiology, vol.100, pp.1-20, 2015.
DOI : 10.1093/treephys/tpv073

URL : https://hal.archives-ouvertes.fr/hal-01557364

J. A. Raven and M. Andrews, Evolution of tree nutrition, Tree Physiology, vol.30, pp.1050-1071, 2010.
DOI : 10.1093/treephys/tpq056

URL : https://academic.oup.com/treephys/article-pdf/30/9/1050/4632147/tpq056.pdf

N. Regier, S. Streb, S. C. Zeeman, and B. Frey, Seasonal changes in starch and sugar content of poplar (Populus deltoides x nigra cv. Dorskamp) and the impact of stem girdling on carbohydrate allocation to roots, Tree Physiology, vol.30, pp.979-987, 2010.

R. R. Reichenbacker, R. C. Schultz, and E. R. Hart, Artificial defoliation effect on Populus growth, biomass production, and total nonstructural carbohydrate concentration, Environmental Entomology, vol.25, pp.632-642, 1996.
DOI : 10.1093/ee/25.3.632

H. Rennenberg, K. Kreutzer, H. Papen, and P. Weber, Consequences of high loads of nitrogen for spruce (Picea abies L.) and beech (Fagus sylvatica L.) forests, New Phytologist, vol.139, pp.71-86, 1998.

H. Rennenberg, W. Seiler, R. Matyssek, A. Gessler, and J. Kreuzwieser, European beech (Fagus sylvatica L.) -a forest tree without future in the south of Central Europe?, Allgemeine Forst Und Jagdzeitung, vol.175, pp.210-224, 2004.

H. Rennenberg, F. Loreto, and A. Polle, Physiological responses of forest trees to heat and drought, Plant Biology, vol.8, pp.556-571, 2006.
DOI : 10.1055/s-2006-924084

H. Rennenberg, M. Dannenmann, A. Gessler, J. Kreuzwieser, J. Simon et al., Nitrogen balance in forest soils: nutritional limitation of plants under climate change stresses, Plant Biology, vol.11, pp.4-23, 2009.
DOI : 10.1111/j.1438-8677.2009.00241.x

C. P. Reyer, N. Brouwers, and A. Rammig, Forest resilience and tipping points at different spatio-temporal scales: approaches and challenges, Journal of Ecology, vol.103, pp.5-15, 2015.
DOI : 10.1111/1365-2745.12337

URL : https://besjournals.onlinelibrary.wiley.com/doi/pdf/10.1111/1365-2745.12337

A. D. Richardson, M. S. Carbone, and T. F. Keenan, Seasonal dynamics and age of stemwood nonstructural carbohydrates in temperate forest trees, New Phytologist, vol.197, pp.850-861, 2013.

J. Roche, M. H. Turnbull, and Q. Guo, Coordinated nitrogen and carbon remobilization for nitrate assimilation in leaf, sheath and root and associated cytokinin signals during early regrowth of Lolium perenne, Annals of Botany, vol.119, pp.1353-1364, 2017.

R. C. Rosecrance, R. S. Johnson, and S. A. Weinbaum, The effect of timing of post-harvest foliar urea sprays on nitrogen absorption and partitioning in peach and nectarine trees, The Journal of Horticultural Science and Biotechnology, vol.73, pp.856-861, 1998.

L. Rowland, A. C. Da-costa, and D. R. Galbraith, Death from drought in tropical forests is triggered by hydraulics not carbon starvation, Nature, vol.528, pp.119-121, 2015.
DOI : 10.1038/nature15539

URL : http://eprints.whiterose.ac.uk/93308/1/Rowland_Nature_paper_accepted_version.pdf

N. K. Ruehr, C. A. Offermann, and A. Gessler, Drought effects on allocation of recent carbon: from beech leaves to soil CO2 efflux, New Phytologist, vol.184, pp.950-961, 2009.

J. Ruffault, N. K. Martin-stpaul, S. Rambal, and F. Mouillot, Differential regional responses in drought length, intensity and timing to recent climate changes in a Mediterranean forested ecosystem, Climatic Change, vol.117, pp.103-117, 2013.

M. G. Ryan, Tree responses to drought, Tree Physiology, vol.31, pp.237-239, 2011.

N. Sade, M. Del-mar-rubio-wilhelmi, K. Umnajkitikorn, and E. Blumwald, Stressinduced senescence and plant tolerance to abiotic stress, Journal of Experimental Botany, vol.69, pp.845-853, 2017.

A. Sakai and W. Larcher, Frost survival of plants. Responses and adaptation to freezing stress, 1987.

A. Sala, F. Piper, and G. Hoch, Physiological mechanisms of drought-induced tree mortality are far from being resolved, New Phytologist, vol.186, pp.274-281, 2010.

A. Sala, D. R. Woodruff, and F. C. Meinzer, Carbon dynamics in trees: Feast or famine?, Tree Physiology, vol.32, pp.764-775, 2012.

S. Salleo, M. A. Lo-gullo, D. De-paoli, and M. Zippo, Xylem recovery from cavitationinduced embolism in young plants of Lauris nobilis: a possible mechanism, New Phytologist, vol.132, pp.47-56, 1996.

Y. Salmon, J. M. Torres-ruiz, R. Poyatos, J. Martinez-vilalta, P. Meir et al., Balancing the risks of hydraulic failure and carbon starvation: a twig scale analysis in declining Scots pine, Plant, Cell and Environment, vol.38, pp.58-72, 2015.

J. J. Sauter and T. Ambrosius, Changes in Partitioning of Carbohydrates in the Wood during Bud Break in Betula pendula Roth, Journal of Plant Physiology, vol.124, pp.31-43, 1986.

J. J. Sauter, B. Vancleve, and S. Wellenkamp, Ultrastrucural and biochemical results on the localization and distribution of storage proteins in a poplar tree and in twigs of the other tree species, Holzforschung, vol.43, pp.1-6, 1989.

J. J. Sauter, Changes in starch content of different parenchyma of trees as induced by temperature, Zeitschrift für Pflanzenphysiologie, vol.56, pp.340-352, 1967.

J. J. Sauter and B. Van-cleve, Seasonal variations of amino acids in the xylem sap of "Populus × canadensis" and its relation to protein body mobilisation, Trees, vol.6, pp.26-32, 1992.

C. Schadel, A. Blochl, A. Richter, and G. Hoch, Short-term dynamics of nonstructural carbohydrates and hemicelluloses in young branches of temperate forest trees during bud break, Tree Physiology, vol.29, pp.901-911, 2009.

K. V. Schäfer, K. L. Clark, N. Skowronski, and E. P. Hamerlynck, Impact of insect defoliation on forest carbon balance as assessed with a canopy assimilation model, Global Change Biology, vol.16, pp.546-560, 2010.

H. J. Schenk and R. B. Jackson, Mapping the global distribution of deep roots in rela-tion to climate and soil characteristics, Geoderma, vol.126, pp.129-140, 2005.

J. Schiebold, M. I. Bidartondo, P. Karasch, B. Gravendeel, and G. Gebauer, You are what you get from your fungi: nitrogen stable isotope patterns in Epipactis species, Annals of Botany, vol.119, pp.1085-1095, 2017.

C. Schimpf and R. Stosser, Histochemische Untersuchungen über die jahreszeitliche Einlagerung und Verteilung von Stärke in Langtrieben beim, Apfel. Mitteilungen Klosterneuburg, vol.34, pp.209-220, 1984.

S. Schmid and S. Palacio, Growth reduction after defoliation is independent of CO 2 supply in deciduous and evergreen young oaks, New Phytologist, vol.214, pp.1479-1490, 2017.

M. Schröter, W. Härdtle, and G. Von-oheimb, Crown plasticity and neighborhood interactions of European beech (Fagus sylvatica L.) in an old-growth forest, European Journal of Forest Research, vol.131, pp.787-798, 2012.

F. Secchi and M. A. Zwieniecki, Patterns of PIP gene expression in Populus trichocarpa during recovery from xylem embolism suggest a major role for the PIP1 aquaporin subfamily as moderators of the refilling process, Plant Cell Environment, vol.33, pp.1285-1297, 2010.

R. Seidl, D. Thom, and M. Kautz, Forest disturbances under climate change, Review, vol.7, pp.395-402, 2016.

R. Seppala, A. Buck, and P. Katila, Adaptation of Forests and People to Climate Change-A Global Assessment Report, IUFRO World Series, vol.22, p.pp, 2009.

R. Serrana, A. Vilagrosa, and J. A. Alloza, Pine mortality in southeast Spain after an extreme dry and warm year: interactions among drought stress, carbohydrates and bark beetle attack, Trees -Structure and Function, vol.29, pp.1791-1804, 2015.

S. Sevanto, N. G. Mcdowell, L. T. Dickman, R. Pangle, and W. T. Pockman, Howdo trees die? A test ofthe hydraulic failure and carbon starvation hypotheses, Plant, Cell & Environment, vol.37, pp.153-161, 2013.

S. Sevanto, Phloem transport and drought, Journal of Experimental Botany, vol.65, pp.1751-1759, 2014.

S. Sevanto and C. Xu, Towards more accurate vegetation mortality predictions, Tree Physiology, vol.36, pp.1191-1195, 2016.

K. K. Shim, J. S. Titus, and W. E. Splittstoesser, The utilization of post-harvest urea sprays by senescing apple leaves, Journal of the American Society for Horticultural Science, vol.97, pp.592-596, 1972.

J. Simon, M. Dannenmann, and R. Gasche, Competition for nitrogen between adult European beech and its offspring is reduced by avoidance strategy, Forest Ecology and Management, vol.262, pp.105-114, 2011.

J. Simon, M. Dannenmann, R. Pena, and A. Gessler, Nitrogen nutrition of beech forests in a changing climate: Importance of plant-soil-microbe water, carbon, and nitrogen interactions, Plant and Soil Marschner Review, vol.1, issue.2, pp.89-114, 2017.

W. Sinclair and G. Hudler, Tree declines: four concepts of causality, Journal of Arboriculture, vol.14, pp.29-35, 1988.

E. W. Sinnott, Factors determining character anddistribution of food reserve in woody plants, Botanical Gazette, vol.66, pp.162-175, 1918.

A. M. Smith and M. Stitt, Coordination of carbon supply and plant growth, Plant, Cell and Environment, vol.30, pp.1126-1149, 2007.

J. Sommer, M. A. Dippold, H. Flessa, and Y. Kuzyakov, Allocation and dynamics of C and N within plant-soil system of ash and beech, Journal of Plant Nutrition and Soil Science, vol.179, pp.376-387, 2016.

T. M. Spann, R. H. Beede, and T. M. Dejong, Seasonal carbohydrate storage and mobilization in bearing and non-bearing pistachio (Pistacia vera) trees, Tree Physiologist, vol.28, pp.207-213, 2008.

D. G. Sprugel, T. M. Hinckley, and W. Schaap, The Theory and Practice of Branch Autonomy, Annual Review of Ecology and Systematics, vol.22, pp.309-334, 1991.

P. E. Staswick, Storage Proteins of Vegetative Plant Tissues, Annual Review of Plant Physiology and Plant Molecular Biology, vol.45, pp.303-322, 1994.

V. Stepien, J. J. Sauter, and F. Martin, Vegetative storage proteins in woody plants, Plant Physiology and Biochemistry, vol.32, pp.185-192, 1994.

M. T. Stevens, E. L. Kruger, and R. L. Lindroth, Variation in tolerance to herbivory is mediated by differences in biomass allocation in aspen, Functional Ecology, vol.22, pp.40-47, 2008.

S. Susiluoto, E. Hilasvuori, and F. Berninger, Testing the growth limi-tation hypothesis for subarctic Scots pine, J Ecol, vol.98, pp.1186-1195, 2010.

Y. Suzuki, A. Makino, and T. Mae, Changes in the turnover of Rubisco and levels of mRNAs of rbcL and rbcS in rice leaves from emergence to senescence, Plant, Cell and Environment, vol.24, pp.1353-1360, 2001.

M. Tagliavini, P. Millard, and M. Quartieri, Storage of foliar-absorbed nitrogen and remobilization for spring growth in young nectarine (Prunus persica var. nectarina) trees, Tree Physiology, vol.18, pp.203-207, 1998.

L. Taiz and E. Zeiger, Plant Physiology, 1998.

W. Tanner and H. Beevers, Transpiration, a prerequisite for long-distance transport of minerals in plants, Plant Biology, vol.98, pp.9443-9447, 2001.

W. Tegel, A. Seim, D. Hakelberg, S. Hoffmann, M. Panev et al., A recent growth increase of European beech (Fagus sylvatica L.) at its Mediterranean distribution limit contradicts drought stress, European Journal of Forest Research, vol.133, pp.61-71, 2014.

E. Teissier-du-cros, F. Le-tacon, and G. Nepveu, Le hêtre, 1981.

F. M. Thomas, R. Blank, and G. Hartmann, Abiotic and biotic factors and their interactions as causes of oak decline in Central Europe, Forest Pathology, vol.32, pp.277-307, 2002.

M. R. Thorpe and P. E. Minchin, Photoassimilate distribution in plants and crops: source-sink relationships, pp.261-282, 1996.

J. Titus and S. M. Kang, Nitrogen metabolism, translocation, and recycling in apple trees, Horticultural Reviews, vol.4, pp.204-246, 1982.

S. Trumbore, Carbon respired by terrestrial ecosystems-recent progress and challenges, Glob Change Biology, vol.12, pp.141-153, 2006.

A. Turcotte, S. Rossi, A. Deslauriers, C. Krause, and H. Morin, Dynamics of depletion and replenishment of water storage in stem and roots of black spruce measured by dendrometers, Frontiers in plant science, vol.2, p.21, 2011.

M. T. Tyree, Desiccation Tolerance of Five Tropical Seedlings in Panama. Relationship to a Field Assessment of Drought Performance, Plant Physiology, vol.132, pp.1439-1447, 2003.

M. Uemura, G. Warren, and P. L. Steponkus, Freezing sensitivity in the sfr4 mutant of Arabidopsis is due to low sugar content and is manifested by loss of osmotic responsiveness, Plant Physiology, vol.131, pp.1800-1807, 2003.

H. Upadhyaya, L. Sahoo, and S. K. Panda, Molecular physiology of osmotic stress in plants, Rout GR, Das AB (ed) Molecular stress physiology of plants, pp.179-192, 2013.

M. Urli, A. J. Porte, H. Cochard, Y. Guengant, R. Burlett et al., Xylem embolism threshold for catastrophic hydraulic failure in angiospermtrees, Tree Physiology, vol.33, pp.672-683, 2013.

M. Uscola, P. Villar-salvador, J. Oliet, and C. R. Warren, Foliar absorption and root translocation of nitrogen from different chemical forms in seedlings of two Mediterranean trees, Environmental and Experimental Botany, vol.104, pp.34-43, 2014.

L. M. Valenzuela-nunez, D. Gerant, P. Maillard, N. Breda, G. G. Cervantes et al.,

L. Valenzuela-nunez, Comparaison interspécifique de la dynamique saisonnière de composes azotes et carbonés chez le chêne sessile (Quercus petraea Matt. Liebl.), le chêne pédonculé (Quercus robur L.) et le hêtre (Fagus sylvatica L.) aux stades juvéniles et adultes, 2006.

, effet de la défoliation et de la lumière sur la gestion des réserves, vol.141

G. W. Van-der-werf, U. Sass-klaassen, and G. M. Mohren, The impact of the 2003 summer drought on the intra-annual growth pattern of beech (Fagus sylvatica L.) and oak (Quercus robur L.) on a dry site in the Netherlands, Dendrochronologia, vol.25, pp.103-112, 2007.

R. Vargas, N. Hasselquist, E. B. Allen, and M. F. Allen, Effects of a hurricane disturbance on aboveground forest structure, arbuscular mycorrhizae and belowground carbon in a restored tropical forest, Ecosystems, vol.13, pp.118-146, 2010.

T. T. Veblen, C. Donoso, T. Kitzberger, and A. J. Rebertus, Ecology of Southern Chilean and Argentinian Nothofagus forests, The ecology and biogeography of Nothofagus forests, pp.293-353, 1996.

M. D. Venturas, J. S. Sperry, and U. G. Hacke, Plant xylem hydraulics: What we understand, current research, and future challenges, Journal of Integrative Plant Biology, vol.59, pp.356-389, 2017.
DOI : 10.1111/jipb.12534

URL : https://onlinelibrary.wiley.com/doi/pdf/10.1111/jipb.12534

L. Vergutz, S. Manzoni, A. Porporato, R. F. Novais, and R. B. Jackson, Global resorption efficiencies and concentrations of carbon and nutrients in leaves of terrestrial plants, Ecological Monographs, vol.82, pp.205-220, 2012.

K. Verheyen, Assessment of the functional role of tree diversity: the multi-site FORBIO experiment, Plant Ecology and Evolution, vol.146, pp.26-35, 2013.

K. Verheyen, M. Vanhellemont, and H. Auge, Contributions of a global network of tree diversity experiments to sustainable forest plantations, Ambio, vol.45, pp.29-41, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01204232

P. Villar-salvador, M. Uscola, and D. F. Jacobs, The role of stored carbohydrates and nitrogen in the growth and stress tolerance of planted forest trees, New Forests, vol.46, pp.813-839, 2015.

P. Vitousek, Nutrient cycling and nutrient use efficiency, The American Naturalist, vol.119, pp.553-572, 1982.
DOI : 10.1086/283931

S. Vizoso, Effets combinés de l'augmentation de la concentration atmosphérique en CO2 et du niveau de fertilisation azotée sur la gestion du carbone et de l'azote chez le chêne pédonculé (Quercus robur) et le hêtre (Fagus sylvatica), p.122, 2004.

J. Voltas, J. J. Camarero, D. Carulla, M. Aguilera, A. Ortiz et al., A retrospective, dual-isotope approach reveals individual predispositions to winter-drought induced tree dieback in the southernmost distribution limit of Scots pine, Plant, Cell & Environment, vol.36, pp.1435-1448, 2013.

S. Wagner, P. Berg, G. Schädler, and H. Kunstmann, High resolution regional climate model simula-tions for Germany: Part II-projected climate changes, Climate Dynamics, vol.40, pp.415-427, 2013.
DOI : 10.1007/s00382-012-1510-1

URL : https://link.springer.com/content/pdf/10.1007%2Fs00382-012-1510-1.pdf

L. A. Wanner and O. Junttila, Cold-induced freezing tolerance in Arabidopsis, Plant Physiology, vol.120, pp.391-399, 1999.
DOI : 10.1104/pp.120.2.391

URL : http://www.plantphysiol.org/content/120/2/391.full.pdf

C. Warren, J. Mcgrath, and M. Adams, Water availability and carbon isotope discrimination in conifers, Oecologia, vol.127, pp.476-486, 2001.
DOI : 10.1007/s004420000609

J. M. Warren, C. M. Iversen, and C. T. Garten, Timing and magni-tude of C partitioning through a young loblolly pine (Pinus taeda L.) stand using 13C labeling and shade treatments, Tree Physiology, vol.32, pp.799-813, 2012.

C. A. Watson, J. M. Ross, U. Bagnares, G. F. Minotta, F. Roffi et al., Environmental-induced modifications to root longevity in Lolium perenne and Trifolium repens, Annals of Botany, vol.85, pp.397-401, 2000.
DOI : 10.1006/anbo.1999.1048

URL : https://academic.oup.com/aob/article-pdf/85/3/397/7983960/850397.pdf

M. A. Watson and B. B. Casper, Morphogenetic constraints on patterns of carbon distribution in plants, Evolution, and Systematics, vol.15, pp.233-258, 1984.

K. L. Webb and J. W. Burley, Stachyose Translocation in Plants, Plant physiology, vol.39, p.973, 1964.
DOI : 10.1104/pp.39.6.973

URL : http://www.plantphysiol.org/content/39/6/973.full.pdf

S. Wetzel, C. Demmers, and J. S. Greenwood, Seasonally fluctuationg bark proteins are a potential form of nitrogen storage in 3 temperate hardwoods, Planta, vol.178, pp.275-281, 1989.

H. Wildhagen, J. Dürr, B. Ehlting, and H. Rennenberg, Seasonal nitrogen cycling in the bark of field-grown Grey poplar is correlated with meteorological factors and gene expression of bark storage proteins, Tree Physiology, vol.30, pp.1096-1110, 2010.

E. Wiley, S. Huepenbecker, B. B. Casper, and B. R. Helliker, The effects of defoliation on carbon allocation: Can carbon limitation reduce growth in favour of storage?, Tree Physiology, vol.33, pp.1216-1228, 2013.

E. Wiley and B. Helliker, A re-evaluation of carbon storage in trees lends greater support for carbon limitation to growth, New Phytologist, vol.195, pp.285-289, 2012.

C. J. Willson, P. S. Manos, and R. B. Jackson, Hydraulic traits are influenced by phylogenetic history in the drought-resistant, invasive genus Juniperus (Cupressaceae), American journal of botany, vol.95, pp.299-314, 2008.

G. Winter, C. D. Todd, M. Trovato, G. Forlani, and D. Funck, Physiological implications of arginine metabolism in plants, Frontiers in Plant Science, vol.6, p.534, 2015.

J. M. Withington, P. B. Reich, J. Oleksyn, and D. M. Eissenstat, Comparisons of structure and life span in roots and leaves among temperate trees, Ecological Monograms, vol.76, pp.381-397, 2006.

C. Zang, C. Hartl-meier, C. Dittmar, A. Rothe, and A. Menzel, Patterns of drought tolerance in major European temperate forest trees: climatic drivers and levels of variability, Glob Change Biology, vol.20, pp.3767-3779, 2014.

B. Zeller, M. Colin-belgrand, É. Dambrine, and F. Martin, 15N partitioning and production of 15N-labelled litter in beech trees following [15N] urea spray, Annals of forest science, vol.55, pp.375-383, 1998.
URL : https://hal.archives-ouvertes.fr/hal-00883208

C. Zhang, S. Meng, Y. Li, L. Su, and Z. Zhao, Nitrogen uptake and allocation in Populus simonii in different seasons supplied with isotopically labeled ammonium or nitrate, TreesStructure and Function, vol.30, pp.2011-2018, 2016.

C. Zhang, G. B. Liu, S. Xue, and C. S. Zhang, Rhizosphere soil microbial properties on abandoned croplands in the Loess Plateau, China during vegetation succession, European Journal of Soil Biology, vol.50, pp.127-136, 2012.

J. Zhao, H. Hartmann, S. Trumbore, W. Ziegler, and Y. Zhang, High temperature causes negative whole-plant carbon balance under mild drought, New Phytologist, vol.200, pp.330-339, 2013.

J. Zimmermann, M. Hauck, . Dulamsuren-ch, and C. Leuschner, Climate warming-related growth decline affects Fagus sylvatica, but not other broad-leaved tree species in Central European mixed forests, Ecosystems, vol.18, pp.560-572, 2015.

R. Zweifel, S. Bangerter, A. Rigling, and F. J. Sterck, Pine and mistletoes: How to live with a leak in the water flow and storage system, Journal of Experimental Botany, vol.63, pp.2565-2578, 2012.